688


0% 20% 40% 60% 80% 100%


Bol Lan Mes Mil Nyc Par Pro


FIGURE 6. Average completeness for the three metrics for each clade. Left red bars (printed black) are the CCMa; middle orange bars (printed light gray) are the CCMb; right brown bars (printed dark gray) are the SCM. Error bars show the SEM. Brackets with stars show the significance of the difference between the two pointed metrics according to Mann-Whitney’s U-test (*, p<0.05; **, p<0.01). Abbreviations used: Bolosauria (Bol), Lanthanosuchoidea (Lan), Mesosauridae (Mes), Millerosauria (Mil), Nycteroleteroidea (Nyc), Pareiasauria (Par), Procolophonoidea (Pro). Color version online.


(51%). MeanCCMbvaries between these peaks and troughs, with a plateau around 58% during the Permian from the Artinskian until the Roadian and a plateau around 66% during the Late Triassic (Carnian, Norian). Mean CCMb increases across the Permian/Triassic boundary. The maximum CCMb of all taxa in each time bin (uppermost gray circles) is again very high with values consistently above 80% and is more stable than the CCMa curve; it is only interrupted in the Ladinian with a mini- mum value of 54%. The minimum (lowermost gray circles) is quite variable, but mostly stays below 40% and in the Wuchiapingian and Changhsingian is even below 10%. There are three time intervals in the early Permian (Assilian–Sakmarian), middle Permian (Wordian–Capitanian), andMiddle/Late Triassic (Ladinian–Carnian) with high minimum values above 50%. The SCM is overall lower than the two CCM


curves (Fig. 3C),with an average mean value of 45%. Amaximummean SCMof 69% is reached in theGhzelian, with a second peak of 59%in the Rhaetian.The overallminimumof 26%occurs in the Anisian.After substantial variation fromthe Sakmarian until the Kungurian, the mean SCM is fairly stable above 40% from the Roadian to the Olenekian across the Permian/Triassic boundary.After this,meanSCM dropsto a trough during the Anisian and Ladinian before rising back to late Permian levels, and thereafter continues to increase throughout the Late Triassic to a Rhaetian peak. As in the CCMa, the


maximumSCMvalue of all taxa in each time bin (uppermost gray circles) is quite stable and high with values above 80% during the entire study interval,with the exception of the troughs in the Sakmarian, Kungurian, and Middle Triassic. The minimum SCM (lowermost gray circles) is very low and consistently under 20%, except during the Sakmarian, Ladinian and Rhaetian. Correlation Tests and Concordance Coefficient.—


Table 2 shows the results of the correlation tests comparing the three completeness metrics and the generic diversity, based on curves with (Table 2A) and without (Table 2B) time bins containing only one genus. In both cases, it appears that none of the completeness metrics are significantly correlated with diversity, using either Spearman’sorKendall’s tests. Including monogeneric stages, comparison of the metrics only shows a highly significant correlation between the SCM and the CCMa for both Spearman’s ρ (ρ=0.671, p=0.006) and Kendall’s τ (τ=0.567, p=0.002) tests. When monogeneric time bins are excluded, the correlation between CCMa and SCM is still significant for both tests: Spearman’s ρ (ρ=0.636, p=0.040) and Kendall’s τ (τ=0.564, p=0.017). In addition, a new significant correlation between CCMa and CCMb is highlighted, but only with Spearman’s ρ (ρ=0.564, p=0.046). Figure 4 presents plotted values of com-


pleteness for the three metrics in pairs. Despite showing an important scatter, the association of CCMa and CCMb on one hand and CCMa and SCM on the other hand seems to follow a


ANTOINE VERRIÈRE ET AL.


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