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792 C. L. Johnson et al.


useful comments. Funding was provided by the Global Environ- mental Facility, as administered through the project Enhancing the Protected Area System of Sulawesi, Fondation Segré and Wildlife Reserves Singapore Conservation Fund.


Author contributions Study design and fieldwork: CJ, RR, LN, HH, AP,WP, IH; data analysis: CJ, FR; writing: CJ, AB, ML, HH, FR.


Conflicts of interest None.


Ethical standards This research abided by the Oryx guidelines on ethical standards.


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Oryx, 2020, 54(6), 784–793 © The Author(s), 2020. Published by Cambridge University Press on behalf of Fauna & Flora International doi:10.1017/S0030605319000851


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