Northern white-cheeked gibbons in Lao 769
FIG. 1 Map of Nam Et-Phou Louey National Protected Area in Lao, showing gibbon survey sites on four management sectors (patrol sectors) and regional ranges of the northern white-cheeked gibbon Nomascus leucogenys.
Gibbons may not perform duets every day and their call-
ing can be affected by factors such as weather conditions and season (Cheyne et al., 2008; Coudrat et al., 2015). We there- fore recorded weather conditions (i.e. presence of direct sunlight, wind, cloud or fog) at the beginning of each survey to assess their possible influence on gibbon detection prob- ability. Weather conditions were recorded as binary values (e.g. any presence of direct sunlight was coded as 1 and complete absence as 0). We recorded five landscape and human disturbance
variables as potential predictors of gibbon group abundance: area of mixed deciduous forest (%), altitude (m) measured at the centre of the sampled area, standard deviation of slope (ruggedness; Riley et al., 1999; Dawrueng et al., 2017), dis- tance (m) from the centre of the sampled area to the bound- ary of the totally protected zone, and a hunting pressure index (Table 1). We selected these variables because gibbon distribution is associated with forest type (Geissmann et al., 2000), altitude and terrain ruggedness (Kim et al., 2011), dis- tance to forest edge or road (Phoonjampa et al., 2011; Akers
et al., 2013), and level of human disturbance (Phoonjampa et al., 2011). We obtained all variables from the databases of the Protected Area headquarters and the records of the WildlifeConservation Society Lao Programme.We calculated
TABLE 1 Description of the site covariates collected and calculated within a 1-km radius around each survey site.
Covariate code
deciduous elevation
slope
distance hunting
Description
Area of mixed deciduous forest (km)
Mean elevation (m)
Standard deviation of slope (ruggedness) (°)
Distance to the boundary of Totally Protected Zone (km)
Hunting pressure index (hunting signs per patrol effort)
Oryx, 2020, 54(6), 767–775 © 2019 Fauna & Flora International doi:10.1017/S0030605318001515 Mean (range) 2.23 (0.16–3.14)
1,369 (1,095–1,978) 9.02 (6.95–11.51)
4.68 (0.00–10.44) 0.04 (0.00–0.24)
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