190


Journal of Paleontology 92(2):189–195


megapixel camera. Images were produced using Zerene Stacker, Zerene Systems LLC, software and were cropped and resized in Photoshop CS6. Anatomical terminology follows the Hymenoptera Anatomy Consortium(http://glossary.hymao.org); wing venation terminology follows Deans and Huben (2003).


Optical calculations.—Ommatidial facet diameters were estimated from the inter-facet center spacing, with facet centers


identified in images that had been pre-processed by high-pass filtering with the Photoshop CS6 unsharp-mask filter, and then segmented using custom software written in Matlab R2015a (Mathworks).


Repository and institutional abbreviation.—The holotype specimen in polished amber (figured here) is deposited in the State Museum of Natural History, Stuttgart (SMNS).


Systematic paleontology Class Insecta Linnaeus, 1758


Order Hymenoptera Fabricius, 1804 Family Evaniidae Latreille, 1802 Genus Parevania Kieffer, 1907


Type species.—Parevania semirufa Kieffer, 1907, by monotypy. Synonyms.—See Deans (2005) for a complete taxonomic history.


Diagnosis.—Eyes large; antennae with 11 flagellomeres; notauli present; fore wing with seven complete cells; fore wing vein 1RS attached to Sc+R basal to pterostigma, curved slightly towards wing base; M+CU often as long as hind jugal lobe; distance between mid- and hind-coxae 0.6 x or less than distance between fore- and mid-coxae; hind tarsomeres not elongated into spines.


Parevania oculiseparata Jennings, Krogmann, and Austin new species Figures 1–3


Holotype.—Male, specimen within a polished piece of amber. Deposited in SMNS (registration SMNS BB-2494).


Occurrence.—An extremely well-preserved specimen as a sin- gle inclusion within Eocene Baltic amber, 56–34 Ma, according to Szwedo and Sontag (2009). Purchased from the collection of Jürgen Velten, Idstein, Germany, by the State Museum of Nat- ural History, Stuttgart.


Diagnosis.—The new species can be diagnosed as a member of the genus Parevania based primarily on the putative synapo- morphy for the genus, fore wing vein 1RS being attached to Sc+R basal to the pterostigma (Fig. 3). Parevania oculiseparata n. sp. differs from all extant species, and from the descriptions of the three (lost) Baltic amber fossil species of Brues (1933), in that the compound eye has two angled surfaces that gives the appearance of having a prominent acute ridge (Fig. 2).


Description (male).—Relatively small compared with most other members of the genus; total body length 4.3mm; head length 0.59mm, mesosoma 1.50mm, metasoma including petiole 2.29mm. Body and metasoma apparently dark brown. Head width 1.07mm; vertex of head 0.5×head width;


compound eye height 0.47mm; eye with prominent acute ridge extending across the entire dorso-ventral elongation of the eye surface (Fig. 2); shortest distance between eyes 0.73mm; distance of lateral ocellus to compound eye 0.17mm; occipital carina obscured; malar space 0.29mm; antennae inserted slightly abovemid-line of eye, distance between toruli 0.85mm; distance from torulus to eye 0.24mm; antenna with 11 flagellomeres; scape 3.97×as long as pedicel; first flagellomere 0.21mm in length; all flagellomeres longer than wide. Mesosoma with surface of mesoscutum and mesoscutellum


rugose-punctate; notauli apparently absent (obscured); propo- deum rugose-punctate; legs elongate; hind leg slightly more robust; hind tibia slightly widened apically; distance between mid- and hind-coxa 0.25 ×distance between fore- and mid- coxae; pretarsal claws short, slender, simple, arolium present. Fore wing venation relatively complete (Fig. 3); first


marginal cell elongate; second submarginal cell separated from first discal cell; 1RS attached to Sc +R basal to pterostigma,


slightly curved; r-rs straight and arising near middle of pterostigma (Fig. 3); pterostigma narrow, ~4 ×as long as wide; 3M and 2CU+3CU nebulous, extending to wing margin; 4RS


Figure 1. Parevania oculiseparata Jennings, Krogmann, and Austin n. sp., holotype ♂ (SMNS BB-2494). (1) Dorsal habitus; (2) ventral habitus. Scale lines=1mm.


Page 1  |  Page 2  |  Page 3  |  Page 4  |  Page 5  |  Page 6  |  Page 7  |  Page 8  |  Page 9  |  Page 10  |  Page 11  |  Page 12  |  Page 13  |  Page 14  |  Page 15  |  Page 16  |  Page 17  |  Page 18  |  Page 19  |  Page 20  |  Page 21  |  Page 22  |  Page 23  |  Page 24  |  Page 25  |  Page 26  |  Page 27  |  Page 28  |  Page 29  |  Page 30  |  Page 31  |  Page 32  |  Page 33  |  Page 34  |  Page 35  |  Page 36  |  Page 37  |  Page 38  |  Page 39  |  Page 40  |  Page 41  |  Page 42  |  Page 43  |  Page 44  |  Page 45  |  Page 46  |  Page 47  |  Page 48  |  Page 49  |  Page 50  |  Page 51  |  Page 52  |  Page 53  |  Page 54  |  Page 55  |  Page 56  |  Page 57  |  Page 58  |  Page 59  |  Page 60  |  Page 61  |  Page 62  |  Page 63  |  Page 64  |  Page 65  |  Page 66  |  Page 67  |  Page 68  |  Page 69  |  Page 70  |  Page 71  |  Page 72  |  Page 73  |  Page 74  |  Page 75  |  Page 76  |  Page 77  |  Page 78  |  Page 79  |  Page 80  |  Page 81  |  Page 82  |  Page 83  |  Page 84  |  Page 85  |  Page 86  |  Page 87  |  Page 88  |  Page 89  |  Page 90  |  Page 91  |  Page 92  |  Page 93  |  Page 94  |  Page 95  |  Page 96  |  Page 97  |  Page 98  |  Page 99  |  Page 100  |  Page 101  |  Page 102  |  Page 103  |  Page 104  |  Page 105  |  Page 106  |  Page 107  |  Page 108  |  Page 109  |  Page 110  |  Page 111  |  Page 112  |  Page 113  |  Page 114  |  Page 115  |  Page 116  |  Page 117  |  Page 118  |  Page 119  |  Page 120  |  Page 121  |  Page 122  |  Page 123  |  Page 124  |  Page 125  |  Page 126  |  Page 127  |  Page 128  |  Page 129  |  Page 130  |  Page 131  |  Page 132  |  Page 133  |  Page 134  |  Page 135  |  Page 136  |  Page 137  |  Page 138  |  Page 139  |  Page 140  |  Page 141  |  Page 142  |  Page 143  |  Page 144  |  Page 145  |  Page 146  |  Page 147  |  Page 148  |  Page 149  |  Page 150  |  Page 151  |  Page 152  |  Page 153  |  Page 154  |  Page 155  |  Page 156  |  Page 157  |  Page 158  |  Page 159  |  Page 160  |  Page 161  |  Page 162  |  Page 163  |  Page 164  |  Page 165  |  Page 166  |  Page 167  |  Page 168  |  Page 169  |  Page 170  |  Page 171  |  Page 172  |  Page 173  |  Page 174  |  Page 175  |  Page 176  |  Page 177  |  Page 178  |  Page 179  |  Page 180  |  Page 181  |  Page 182  |  Page 183  |  Page 184  |  Page 185  |  Page 186  |  Page 187  |  Page 188  |  Page 189  |  Page 190  |  Page 191  |  Page 192  |  Page 193  |  Page 194  |  Page 195  |  Page 196  |  Page 197  |  Page 198  |  Page 199  |  Page 200  |  Page 201  |  Page 202  |  Page 203  |  Page 204