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Signorelli and Raven—Current knowledge of the family Cardiliidae


Mactroidea is unconfirmed, but retained. Besides type material of described species, scarce additional specimens are available in malacological collections. The position of Cardilia between Mactridae and Myidae, mentioned by Marwick (1943), was never tested. Currently, the superfamily Mactroidea includes four families (Signorelli and Carter, 2016). They are Mactridae, which originated in the Late Cretaceous, Anatinellidae, Meso- desmatidae, and Cardiliidae, all recorded from the Eocene. All families are characterized as shallow or deep burrowers, occurring in the intertidal zone or on the shelf. This work is part of an ongoing revision of superfamily Mac-


troidea, including all recent and fossil species. This study includes morphological data both fromliterature and from new observations of type material and non-type specimens, and states the basis for future studies on recent and fossil species of Cardiliidae.


Conclusions


The Cardiliidae comprise a single genus, Cardilia, which is known from middle Eocene to Recent. Most species are rare and material available is very limited. Eleven extinct species (including one that is described in this paper) and four extant species (for which fossil material is remarkably scarce) are recognized. The placement of the family Cardiliidae in the superfamily Mactroidea requires confirmation.


Acknowledgments


Special thanks to V. Herós and P. Maestrati (MNHN); A. Salvador, K. Way, and J. Todd (NHMUK); T. Schiøtte (ZMUC); B. Hoeksema, F. Wesselingh, and R. Pouwer (Naturalis); and M. Harzhauser and O. Mandic (NHMW) for their assistance in the revision of type material and additional examined speci- mens. Many thanks to G. Rosenberg and S. Popov for their comments. JHS acknowledges CONICET of Argentina. This work was partially supported by the PICT 640 of the Agencia Nacional de Investigaciones Científicas y Técnicas. This is publication N° 89 of the Laboratorio de Reproducción y Biología Integrativa de Invertebrados Marinos (LARBIM).


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