218


Journal of Paleontology 92(2):207–220


identified throughout the Tethys region, including Poland (Narkiewicz, 1999), Romania (Orchard et al., 2007a), Turkey (Nicora, 1977), China (Orchard et al., 2007b), India (Chhabra and Kumar, 1992), and Tibet (Wang and Wang, 1976; Tian, 1982). It is not clear from the published illustrations whether the Tethyan specimens represent species that can be identified in North America, so detailed correlation between these regions is not pos- sible at this time. However, the occurrence of Magnigondolella n. gen. close to the proposed position of the Olenekian-Anisian boundary in the sections at Desli Caira in Romania (Orchard et al., 2007a) and Guandao in China (Orchard et al., 2007b) suggests that the genus is potentially important for the recognition of this boundary. Further study of specimens of Magnigondolella n. gen. from these sections will be necessary as work on defining and correlating the Olenekian-Anisian boundary continues. The recognition of five new species of Magnigondolella


Additional specimens belonging to the genus can also be


n. gen., and three additional species in open nomenclature, increases the number of conodont species identified from the lower and middle Anisian of North America considerably. The early and middle parts of the Anisian were characterized as a low diversity period globally by Chen et al. (2016) and Kiliç et al. (2016), with both studies recognizing ~30 species for the Aegean, Bithynian, and Pelsonian (equivalent to the lower and middle Anisian). Such a low value may be due partially to the preliminary nature of the study of North American Anisian taxa, and partially to endemism of North American species. This highlights the need to carry out detailed taxonomic work before accurate judgments of biodiversity can be made.


Acknowledgments


The authors wish to thank Associate Editor H.-D. Sues and an anonymous reviewer for comments that helped to improve this manuscript. The samples from B.C. utilized in this study were collected by the authors as well as by E.T. Tozer and J.-P. Zonneveld. The samples from Nevada were largely derived from the matrix of ammonoid collections made by H. Bucher. Samples were processed and specimens photographed by P. Krauss and H. Taylor. Financial support for this study was provided by the Cordillera Project of the Geological Survey of Canada GEM-2 Program.


References


Bender, H., 1970, Zur Gliederung der Mediterranen Trias II. Die Conodontenchronologie der Mediterranen Trias: Annales Géologiques des Pays Helléniques, v. 19, p. 465–550.


Bucher, H., 1988, A new Middle Anisian (Middle Triassic) ammonoid zone from northwestern Nevada (USA): Eclogae Geologicae Helvetiae, v. 81, p. 723–762.


Bucher, H., 1989, Lower Anisian ammonoids from the northern Humboldt Range (northwestern Nevada, USA) and their bearing upon the Lower-Middle Triassic boundary: Eclogae Geologicae Helvetiae, v. 82, p. 945–1002.


Bucher, H., 1992a, Ammonoids of the Hyatti Zone and the Anisian transgression in the Star Peak Group, northwestern Nevada, USA: Palaeontographica Abteilung A, v. 223, p. 137–166.


Bucher, H., 1992b, Ammonoids of the Shoshonensis Zone (Middle Anisian, Middle Triassic) from northwestern Nevada (USA): Jahrbuch der Geolo- gischen Bundesanstalt Wien, v. 135, p. 425–465.


Bucher, H., 1994, New ammonoids from the Taylori Zone (Middle Anisian, Middle Triassic) from northwestern Nevada (USA): Mémoires de Géologie (Lausanne), v. 22, p. 1–8.


Table 2. Table showing the number of P1 elements of Magnigondolella n. gen. species identified from localities in B.C. and Nevada. British Columbia


lower Anisian M. alexanderi M. regalis 22 4 7 —— — 1 — middle Anisian 11 5 5 3 M. cyri 1632 4


M. salomae ——— — — — Total


23


— 10


3 4 12 lower Anisian M. alexanderi 10 1 6 2 M. cyri —— 510


65 34


78 6 1 11 21 —— 13 — 21 — 18


M. n. sp. A ——— — 21 — ——— — M. n. sp. B ——— —


M. n. sp. C ——— — — Total


10 1 16 20


M. salomae ——— — 21 —— 2 5 ——— — 131


4 114 10 26 1 49


43 3


8 8


13 7


— 4 —— 7


M. julii ——— — — — 32 —— — M. nebuchadnezzari ——— — —


7 3 1 8 2 Nevada middle Anisian


C-176327 C-176347 C-176338 C-159809 C-176314 C-176324 C-201551 C-176339 C-176321 C-176337 C-176317 C-176325 C-175807 C-175803 C-175805 C-175806 C-175804 C-176526 —


—


M. julii ——— 84 11 —— — 210 M. nebuchadnezzari —— 5


4 —— — —


7 5


—


8 8


2 6


1 1


3 13


— — ————— — — —


— — — — ————— — — 5


22 4 14 20 8 2 1 —— —


— ————— — — 26


Total 265


134


2 ———— — 441 — 42


3 —— 223 5 4 6


22 22 500


— 32 11 —


— —— ——— — — — 2 ——— — 9


1 —— 11 — 21 — — — 15


17 —


2 1


1 7 upper Anisian


C-201922 O-99574 O-42349 C-209952 V-002992 V-002995 C-302187 V-002955 V-002957 V-002999 V-002506 C-101077 O-68294 O-99575 O-99588 C-090852 V-002606 V-002978 V-002979 V-002980 Total — 27


4


8 —— — 5


3 4 — 2


1 ——— — — — — 2 ——— — 21


2 4 4 83


— ——— 44 —


— 215


— 11 1 —— 14 1


1 —— 5 9


162


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