486
Journal of Paleontology 92(3):478–487
have been able to colonize land across long distances of open ocean. A premier example is the colonization of the Hawaiian Islands by various millipedes (Shelley and Golovatch, 2011, p. 33, 68), presumably by rafting. The paucity of millipedes in the Luoping biota contrasts with the more common occurrence of millipedes in some assemblages with a nonmarine origin, for instance in the Cretaceous site of Las Hoyas, Spain, where millipedes are more numerous, but still only a small proportion of the fauna (Selden and Shear, 2016).
Acknowledgments
This work was supported by the National Natural Science Founda- tion of China (nos. 41502013, 41772022 and 41661134047), Chinese Geological Survey projects (Grant nos. DD20160020, 12120114068001, 1212011140051, 1212011120621, and 1212010610211). JTH thanks colleagues at the Paleontological Institute, Moscow, and the National Museum of the Czech Republic, for allowing access to material from Russia and Bohemia. We also thank two anonymous reviewers for their insightful comments.
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