Nardin et al.—Transitional blastozoan from the middle Cambrian of the Czech Republic


673


of the Paradoxides (P.) paradoxissimus gracilis trilobite Zone. The holotype SZ349 was collected about 10mabove the base of the Paradoxides (P.) paradoxissimus gracilis Biozone at the Jince-Vinice locality. The Paradoxides (P.) paradoxissimus gracilis Biozone is correlated with the middle and higher levels of the Baltic Paradoxides (P.) paradoxissimus Biozone Biozone (Fatka et al., 2014). This level corresponds to the Drumian, Cambrian Series 3 in the new international global stratigraphic chart (Babcock et al., 2004; Geyer and Landing, 2004; Fatka, 2006; Zamora et al., 2009; Fatka et al., 2014).


Materials


Repositories and institutional abbreviations.—Specimens of Felbabkacystis n. gen. and comparative materials of Vyscystis ubaghsi Fatka and Kordule, 1990 are deposited in the collections of the Czech Geological Survey, Prague (SZ, LK, MI) and the National Museum, Prague (NML). Comparative materials of Lepidocystis wanneri Foerste, 1938 and Kinzercystis durhami Sprinkle, 1973 are deposited in the Museum of Comparative Zoology, Harvard University, Cambridge,Massachusetts (MCZ).


Systematic paleontology Subphylum Blastozoa Sprinkle, 1973


Figure 1. (1) Geographical location of fossiliferous sites in the Příbram- Jince Basin. 1 = Rejkovice-Ostrý Hill; 2 = Hill slope of Vinice near Jince. (2) Synthetic stratigraphic column of the Jince Formation (Stage 5–Drumian) in the Příbram-Jince Basin with the stratigraphic range of the echinoderm species. Modified from Fatka and Szabad (2014) and Fatka et al. (2015).


can occur abundantly but in restricted areas. Better-preserved echinoderm material is composed of fully articulated specimens either isolated or clustered. All known specimens of Felbabkacystis luckae n. gen.


n. sp. were collected in two fossil sites: (1) Rejkovice-Ostrý Hill locality (49.8207ºN, 13.9593ºE) (samples SZ343, SZ349, MI2, LK1-2); and (2) Jince-slope called Jince-Vinice locality (49.7844ºN, 13.9930ºE) (samples SZ344-347) (Fig. 1.1). Both are situated in the Jince Formation, within the lower third


Remarks.—The subphylum Blastozoa contains at present ten classes. Since Chauvel (1941), many authors considered that the class Eocrinoidea “includes a heterogeneous assemblage of species whose only similarity is that they lack the autapo- morphic characteristics of the other, less ambiguously defined, cystoid [blastozoan] groups” (Smith, 1984, p. 439). Eocrinoids are mostly defined as basal blastozoans (brachiole-bearing echinoderms) having an irregular plated body wall (with imbricate or tessellate plating), with or without epispires, and an irregularly multiplated stalk or a holomeric stem (Ubaghs, 1968; Sprinkle, 1973; Broadhead, 1982; Paul and Smith, 1984). “The diagnoses [of the class Eocrinoidea] are so broad that they essentially define those blastozoans that are not clearly assign- able, by default, to a cystoid or blastoid group” (Parsley and Zhao, 2006, p. 1063). Therefore, the class Eocrinoidea, which corresponds to a paraphyletic assemblage, is not considered here as a valid taxonomic entity, and basal-most blastozoans are not assigned to any existing class.


Family Lepidocystoidae Durham, 1968 Figure 2


Diagnosis.—‘Calyx-bearing’ blastozoans. Oral disc made of numerous adjacent plates, with simple sutural pores along their sutures; anal pyramid, hydropore-gonopore located in the ‘CD’ interradius in the oral disc. Ambulacral system confined to oral disc, consisting of a central, covered mouth, several radially arranged ambulacral grooves embedded into the oral surface, and numerous straight or coiled biserial brachioles alternately arranged alongside each ambulacral groove. Aboral cup cone- shaped or elongated into a cylindrical stalk, and composed of numerous imbricate plates.


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