Cole et al.—New crinoids from the Middle and Upper Ordovician, Spain


Material.—The taxon is known from a single specimen, MPZ2016/104.


Occurrence.—Fombuena Formation, Huerva Member, upper middle Berounian (lower Katian, Upper Ordovician), near Fombuena (Zaragoza province), Spain; Locality 2.


Camerata genus indeterminate species indeterminate Figure 6.2


Remarks.—This taxon is represented by a single partial calyx. The specimen consists of the proximal portion of a crushed calyx with proximal stem attached. Calyx plates are large, all of approximately similar size, flat to slightly convex, lacking median ray ridges or ornamentation. The stem is circular, heteromorphic, with a large circular lumen. One partial interray with plating 1-3-?; the first interradial plate hexagonal, distally tapered, separating adjoining radials. The lack of any plate ornamentation and the relative sizes


of the calyx plates indicate this specimen represents a distinct species from this fauna. The poor preservation and degree of disarticulation of the specimen prevents confident identification of other circlet plates and precludes assignment beyond that of subclass.


Material.—The taxon is known from a single specimen, MPZ2016/93.


Occurrence.—Fombuena Formation, Huerva Member, upper middle Berounian (lower Katian, Upper Ordovician), near Fombuena (Zaragoza province), Spain; Locality 2.


Order Monobathrida Moore and Laudon, 1943 Suborder Glyptocrinina Moore, 1952


Superfamily Patelliocrinoidea Angelin, 1878 Family Patelliocrinidae Angelin, 1878 Genus Eopatelliocrinus Brower, 1973


Type species.—Eopatelliocrinus scyphogracilis Brower, 1973; by original designation.


Other species.—E. latibrachiatus Brower, 1973; E. ornatus (Billings, 1857); E. hispaniensis n. sp.


Occurrence.—Previously, Eopatelliocrinus was known exclu- sively from the Katian of North America (Brower, 1973, 1994). The occurrence of E. hispaniensis in the Katian of Spain expands its known range to Laurentia and Gondwana.


Remarks.—The occurrence of Eopatelliocrinus in Spain represents only the third crinoid genus to occur in both Gond- wana and Laurentia during the Ordovician (Zamora et al., 2015b). The two other examples are the disparid Iocrinus Hall, 1866 and the presumed cladid Merocrinus Walcott, 1883. These two genera are relatively long ranging and known from numerous localities, which makes the appearance of Eopatelliocrinus in Gondwana somewhat unexpected and highlights the utility of improved sampling from under- represented paleogeographic regions.


Eopatelliocrinus hispaniensis new species Figures 4.7, 5.7


2015a camerate crinoid Zamora et al., p. 237, fig. 19F.


Holotype.—The holotype, MPZ2016/81, is the only known specimen of this species.


Diagnosis.—Calyx relatively small, medium cone shaped; basal rim present; smooth plate sculpturing, no depression at calyx plate boundaries; ray ridges extend onto fixed brachials; arms robust, taper gently; proximal column circular.


Occurrence.—Fombuena Formation, Huerva Member, upper middle Berounian (lower Katian, Upper Ordovician), near Fombuena (Zaragoza province), Spain; Locality 2.


Description.—Calyx large in size; medium vase shaped, base truncate; calyx plates gently convex or concave, smooth sculp- turing, strongly convex along ray plates. Basal circlet visible in side view, truncate proximally with


proximal ridge, approximately 19% of calyx height; basal plates concave, total number unknown. Radial circlet approximately 24% of calyx height, radial


plates presumably five, hexagonal, as wide as high. Radial facets with fixed first primibrachials, but approximately 40% of distal width of radial plate and declivate. Normal interrays very poorly preserved, depressed


between ray ridges, in contact with tegmen. Presumably one large first interradial plate followed by two interradial plates; distal-most fixed interradial plates adjacent to first or second secundibrachial. Posterior interray and tegmen unknown. First primibrachial fixed, tetragonal, approximately as wide as high, much smaller than radial plates; second


primibrachial fixed, axillary, pentagonal. First secundibrachial fixed, tetragonal; second secundibrachial tetragonal, at least partially fixed. Small fixed intrabrachial plates in center of ray begin adjacent to secundibrachials but not in contact with primaxil. Free arms 10, atomous. Third secundibrachial free, cuneate


uniserial, bearing stout pinnule abaxially. Remainder of free brachials cuneate uniserial, aborally convex, pinnulate. Pinnules long, stout. Column circular, holomeric, heteromorphic. In proxistele,


columnal latera convex, various numbers of internodals. Column fully developed in mesistele with pattern of N3231323; epifacet of nodals greatly expanded such that nodal diameter twice the diameter of internodals; internodal columnals with convex latera. Lumen and columnal facets unknown.


Etymology.—Hispaniensis, in reference to the Spanish origin of the specimen.


Measurements.—Holotype: crown height, 13.6; calyx height, 3.9; calyx maximum width, 3.7; proximal basal circlet width, 1.2; basal circlet height, 0.95; radial circlet height, 1.4; column


729


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