Allaire et al.—Early Ordovician eocrinoids from Morocco
Animal Life’), coupled with cladistic and morphometric ana- lyses, made it possible to reassess the morphological variability and thus to propose a critical revision of this genus. Rhopalo- cystis now appears to be composed of only five morphological species: R. destombesi, R. fraga, R. grandis, R. havliceki, and R. zagoraensis. Considering the systematic revisions proposed here,
R. dehirensis, R. lehmani, and the three species left in open nomenclature (R. sp. A, R. sp. B, and R. sp. C) should be now considered as invalid. The specimens originally identified as R. dehirensis (Chauvel and Régnault, 1986) are attributed to R. destombesi; those identified as R. lehmani (Chauvel and Régnault, 1986) are assigned to R. zagoraensis. The specimens corresponding to R. sp. A and R. sp. B (Chauvel, 1971, 1978) are assigned to R. fraga,and R. sp. C (Chauvel, 1978) is now identified as R. havliceki. The polymorphismof some species, the presence of growth allometry, and more generally the very large morphological disparity characterizing the genus Rhopalocystis probably explain the previously overestimated intrageneric specificrichness.
Acknowledgments
This study was part of the project RALI (Rise of Animal Life), funded by theANR (Agence Nationale de la Recherche, France; grant number ANR-11-BS56-0025), and part of the CNRS- CNRST cooperation program VALORIZ (grant number 52943). The authors are particularly thankful to M. Nohejlová (Charles University, Prague, Czech Republic), B. Deline (University of West Georgia, Carrollton, United States), and I.A. Rahman (University of Oxford, Oxford, England) for reviewing the manuscript and making many helpful remarks. The authors are also grateful to D. Berthet (Musée des Confluences, Lyon, France), K. El Hariri (Université Cadi Ayyad, Marrakesh, Morocco), F.J. Lindemann and H.A. Nakrem (Natural History Museum, Oslo, Norway), J. Plaine (Institut de Géologie, Rennes, France), A. Prieur and E. Robert (Université Lyon 1, Villeurbanne, France), and S. Régnault (Muséum d’Histoire naturelle, Nantes, France) for providing access to specimens and/or latex casts of specimens housed in their collections. Precious field assistance was provided by A. Bachnou and A. Hafid (Marrakesh), B. Pittet and H. Hormière (Villeurbanne), M. Masrour (Agadir), F. Noailles (Bristol), H. Nowak and T. Vandenbroucke (Villeneuve d’Ascq), R. and V. Reboul (Saint-Chinian), M. Vidal (Brest), and D. Vizcaïno (Carcassonne). The authors also thank mad meg, the artist who made the beautiful illustration of Rhopalocystis destombesi (Fig. 2.5).
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