Cole et al.—New crinoids from the Middle and Upper Ordovician, Spain
721
Table 2. Measurements for diplobathrid crinoid specimens. All measurements are in millimeters. Asterisk (*) indicates measurement of compressed or incom- plete specimen; double asterisk (**) indicates holotype specimen; question mark (?) indicates feature not measured due to nonpreservation.
Taxon
Fombuenacrinus nodulus
Goyacrinus gutierrezi MPZ 2016/99** Ambonacrinus decorus MPZ 2016/94**
Fombuenacrinus nodulus
Fombuenacrinus nodulus
Fombuenacrinus nodulus
Specimen number
Crown height
MPZ 2016/105** 34.7* MPZ 2016/97 MPZ 2016/84 MPZ 2016/92
6.5 ?
28.1* 22.6
29.3
Dalicrinus hammanni MPZ 2016/103** 56.4* Dalicrinus hammanni MPZ 2016/100
27.7*
Calyx height
10.2 1.8 6.9
11.9* 5.2
5.5
13.7* 16.0*
Anthracocrinidae (fixed pinnules and lack of anitaxial plating in the posterior interray) are no longer invariable in all genera assigned to the family. In particular,Goyacrinus gutierrezi n. gen. n. sp. possesses an anitaxial ridge in the posterior interray and lacks fixed pinnules, and Fombuenacrinus nodulus n. gen. n. sp. variably possesses fixed pinnules. Although these two new taxa differ in their possession of these characters, they are still unambi- guously unitedwith other genera assigned toAnthracocrinidae on the basis of the characters given in the preceding emended diag- nosis. Because these traits represent a unique suite of characters among Ordovician diplobathrids, emending the diagnosis of Anthracocrinidae has been deemed amore conservative approach than erecting a new family to hold these genera. The genera previously assigned to Anthracocrinidae are
known only from the Ordovician of Laurentia. Thus, the addition of three new genera extends the known geographic range of Anthracocrinidae to Gondwana.
Genus Fombuenacrinus new genus
Type species.—Fombuenacrinus nodulus new species, by monotypy.
Diagnosis.—As for species, by monotypy.
Etymology.—Fombuenacrinus is derived from the village of Fombuena, which is near the site of collection.
Remarks.—Fombuenacrinus is assigned to the Anthracocrinidae on the basis of confinement of the infrabasals and proximal basals to the basal concavity, radials separated by a large first interray plate, fixed brachials bifurcating twice within the calyx giving rise to secundibrachials and tertibrachials, 20 free arm openings, lower pinnules enlarged and variably fixed, and undifferentiated posterior interray. Fombuenacrinus differs from most Anthracocrinidae in that only the lowest pinnule is variably fixed. All other genera
previously assigned to Anthracocrinidae possess numerous fixed pinnules in rays, half rays, and often quarter rays. Fombuenacrinus is most similar to Anthracocrinus Strimple and Watkins, 1955 but is differentiated by a globose to bowl-shaped calyx, highly convex median ray ridges, interrays in contact with the tegmen, few fixed pinnules, 20 free arms, and biserial brachials. By contrast, Anthracocrinus has a conical to
Type material.—The holotype (MPZ2016/105) and four para- types (MPZ2016/84, 85, 92 [part and counterpart], 97).
Diagnosis.—Anthracocrinid with globose to bowl-shaped calyx; highly convex median ray ridges; infrabasals confined to basal concavity; first interray plate very large; all interrays contiguous with the tegmen; proximal interray plating 1–2?; CD
interray not differentiated from other rays; four arms per ray; fixed secundibrachials and tertibrachials; intrabrachial plates between fixed secundibrachials and tertibrachials of adjacent half rays and between fixed tertibrachials of adjacent quarter rays; free arms unbranched; first pinnule variably fixed; first two pinnules enlarged; mature brachials wedge biserial.
Occurrence.—Fombuena Formation, Huerva Member, upper middle Berounian (lower Katian, Upper Ordovician), near Fombuena (Zaragoza province), Spain; Localities 1 and 2.
Description.—Calyx globose to bowl shaped, wider than high;
plates with fine stellate ornamentation; plate sutures indistinct, flush with plate surface; shallow basal concavity surrounded by continuous pentagonal rim; ray ridges distinct, bulbous, widest at plate sutures, ornamented with parallel ridges; basals and radials ornamented with heavy ridges forming a pentastellate pattern at base of calyx.
Max. calyx width
15.1* 2.9
11.2
21.4* 9.9*
7.3* 21.7* 11.04*
Proximal basal circlet width
7.24* 1.5 4.4
?
4.5* 2.5
? ?
Basal circlet height
2.7 0.5 2.6 ?
1.4 1.9 ? ?
Radial circlet height
2.6 0.8 2.7 3.4 1.9
3.2
1.2 ?
subconical calyx, broad and low median ray ridges, fixed pinnules that prevent contact of interrays with the tegmen, 15 free arms, and uniserial brachials. Fombuenacrinus also shares many similarities with Visocrinus Ausich et al., 2002 from the Katian of Spain. Visocrinus is currently assigned to Rhodocrinitidae but possesses most diagnostic features of Anthracocrinidae. Thus, family reassignment may be in order for this genus but is beyond the scope of this study. Fombuenacrinus is differentiated from Visocrinus by its undifferentiated posterior interray, large first interradial plate, and brachials fixed to the third tertibrachial. By contrast, Visocrinus has a differentiated posterior interray, small first interradial plate, and brachials fixed to the fourth tertibrachial.
Fombuenacrinus nodulus new species Figures 3.1, 3.2, 3.7, 3.8, 3.11, 3.12, 4.1, 4.2
2015a camerate crinoid in Zamora et al., p. 237, figs. 19K, 19L
Page 1 |
Page 2 |
Page 3 |
Page 4 |
Page 5 |
Page 6 |
Page 7 |
Page 8 |
Page 9 |
Page 10 |
Page 11 |
Page 12 |
Page 13 |
Page 14 |
Page 15 |
Page 16 |
Page 17 |
Page 18 |
Page 19 |
Page 20 |
Page 21 |
Page 22 |
Page 23 |
Page 24 |
Page 25 |
Page 26 |
Page 27 |
Page 28 |
Page 29 |
Page 30 |
Page 31 |
Page 32 |
Page 33 |
Page 34 |
Page 35 |
Page 36 |
Page 37 |
Page 38 |
Page 39 |
Page 40 |
Page 41 |
Page 42 |
Page 43 |
Page 44 |
Page 45 |
Page 46 |
Page 47 |
Page 48 |
Page 49 |
Page 50 |
Page 51 |
Page 52 |
Page 53 |
Page 54 |
Page 55 |
Page 56 |
Page 57 |
Page 58 |
Page 59 |
Page 60 |
Page 61 |
Page 62 |
Page 63 |
Page 64 |
Page 65 |
Page 66 |
Page 67 |
Page 68 |
Page 69 |
Page 70 |
Page 71 |
Page 72 |
Page 73 |
Page 74 |
Page 75 |
Page 76 |
Page 77 |
Page 78 |
Page 79 |
Page 80 |
Page 81 |
Page 82 |
Page 83 |
Page 84 |
Page 85 |
Page 86 |
Page 87 |
Page 88 |
Page 89 |
Page 90 |
Page 91 |
Page 92 |
Page 93 |
Page 94 |
Page 95 |
Page 96 |
Page 97 |
Page 98 |
Page 99 |
Page 100 |
Page 101 |
Page 102 |
Page 103 |
Page 104 |
Page 105 |
Page 106 |
Page 107 |
Page 108 |
Page 109 |
Page 110 |
Page 111 |
Page 112 |
Page 113 |
Page 114 |
Page 115 |
Page 116 |
Page 117 |
Page 118 |
Page 119 |
Page 120 |
Page 121 |
Page 122 |
Page 123 |
Page 124 |
Page 125 |
Page 126 |
Page 127 |
Page 128 |
Page 129 |
Page 130 |
Page 131 |
Page 132 |
Page 133 |
Page 134 |
Page 135 |
Page 136 |
Page 137 |
Page 138 |
Page 139 |
Page 140 |
Page 141 |
Page 142 |
Page 143 |
Page 144 |
Page 145 |
Page 146 |
Page 147 |
Page 148 |
Page 149 |
Page 150 |
Page 151 |
Page 152 |
Page 153 |
Page 154 |
Page 155 |
Page 156 |
Page 157 |
Page 158 |
Page 159 |
Page 160 |
Page 161 |
Page 162 |
Page 163 |
Page 164 |
Page 165 |
Page 166 |
Page 167 |
Page 168 |
Page 169 |
Page 170 |
Page 171 |
Page 172 |
Page 173 |
Page 174 |
Page 175 |
Page 176 |
Page 177 |
Page 178 |
Page 179 |
Page 180 |
Page 181 |
Page 182 |
Page 183 |
Page 184 |
Page 185 |
Page 186 |
Page 187 |
Page 188 |
Page 189 |
Page 190 |
Page 191 |
Page 192 |
Page 193 |
Page 194 |
Page 195 |
Page 196 |
Page 197 |
Page 198 |
Page 199 |
Page 200 |
Page 201 |
Page 202 |
Page 203 |
Page 204 |
Page 205 |
Page 206 |
Page 207 |
Page 208 |
Page 209 |
Page 210 |
Page 211 |
Page 212 |
Page 213 |
Page 214 |
Page 215 |
Page 216 |
Page 217 |
Page 218 |
Page 219 |
Page 220 |
Page 221 |
Page 222 |
Page 223 |
Page 224 |
Page 225 |
Page 226 |
Page 227 |
Page 228 |
Page 229 |
Page 230 |
Page 231 |
Page 232 |
Page 233 |
Page 234 |
Page 235 |
Page 236 |
Page 237 |
Page 238 |
Page 239 |
Page 240 |
Page 241 |
Page 242 |
Page 243 |
Page 244 |
Page 245 |
Page 246 |
Page 247 |
Page 248 |
Page 249 |
Page 250 |
Page 251 |
Page 252 |
Page 253 |
Page 254 |
Page 255 |
Page 256 |
Page 257 |
Page 258 |
Page 259 |
Page 260 |
Page 261 |
Page 262 |
Page 263 |
Page 264 |
Page 265 |
Page 266 |
Page 267 |
Page 268 |
Page 269 |
Page 270 |
Page 271 |
Page 272 |
Page 273 |
Page 274 |
Page 275 |
Page 276 |
Page 277 |
Page 278 |
Page 279 |
Page 280 |
Page 281 |
Page 282 |
Page 283 |
Page 284 |
Page 285 |
Page 286 |
Page 287 |
Page 288
