730


Journal of Paleontology 91(4):715–734


height, 7.4* (measurements in millimeters, *incomplete or specimen crushed).


Remarks.—The relatively small calyx, three plates above the primanal, relatively numerous fixed brachials, three unequal basal plates, and 10 cuneate arms with prominent pinnules ally this taxon with Eopatelliocrinus. Species characters in Eopatelliocrinus include shape of the calyx, relative size, pre- sence or absence of a proximal rim on the basal circlet, calyx plate sculpturing, presence or absence of ray ridge extending onto fixed brachials, nature of arms, and shape of proximal column. E. hispaniensis n. sp. is distinct from other species in this genus with a medium cone-shaped calyx, the calyx relatively small, a rim on the proximal basal circlet, smooth calyx plate sculpturing with plate triple junctions not depressed, ray ridges that continue onto fixed brachials, robust and gently tapering arms, and a circular proximal column. By contrast, E. latibrachiatus Brower, 1973 has a medium globe-shaped calyx, the calyx relatively small, rim absent on the proximal basal circlet, broadly stellate plate sculpturing with plate triple junctions depressed, ray ridges that do not continue onto fixed brachials, slender arms that taper sharply in distal third, and a weakly pentalobate proximal column; E. ornatus Brower, 1994 has a medium bowl-shaped calyx, the calyx relatively small, a rim on the proximal basal circlet, broad and fine stellate calyx plate sculpturing, plate triple junctions not depressed, ray ridges that continue onto fixed brachials, slender and gently tapering arms, and a circular distal column; and E. scyphogracilis has a medium cone-shaped calyx, the calyx relatively large, a rim on the proximal basal circlet, weak stellate calyx plate sculpturing,


plate triple junctions not depressed, ray ridges that continue onto fixed brachials, slender and gently tapering arms, and a circular distal column.


Subclass Cladida Moore and Laudon, 1943 Order Dendrocrinida Bather, 1899


Superfamily Mastigocrinoidea Bather, 1899 Family Thenarocrinidae Jaekel, 1918 Genus Picassocrinus new genus


Type species.—Picassocrinus villasi new species, by monotypy.


Diagnosis.—Thenarocrinid with relatively small crown, three anal plates in sutural contact above the anal X, arms branch twice isotomously, and brachials as high as wide.


Etymology.—Picassocrinus, in recognition of the Spanish abstract artist Pablo Picasso and in reference to the atypical plating of the posterior interray.


Picassocrinus villasi new species Figures 4.8, 5.8–5.10


2015a cladid crinoid in Zamora, et al., p. 237, fig. 19B.


Holotype.—The holotype, MPZ2016/83, part and counterpart, is the only known specimen of this species. Diagnosis.—As for genus, by monotypy.


Occurrence.—Fombuena Formation, upper middle Berounian (lower Katian, Upper Ordovician), near Fombuena (Zaragoza province), Spain; Locality 2.


Description.—Crown small; aboral cup medium cone shaped, width to height ratio approximately 1.0, plates gently convex, smooth.


Infrabasals five, equal in size, approximately as high as


wide; infrabasal circlet approximately 28% of aboral cup height, base truncate. Basals probably five, hexagonal, approximately as high as wide, approximately the same size as radial plates;


basal circlet approximately 34% of aboral cup height. Radials five, pentagonal, markedly convex distally emphasizing radial position on plate, wider than high; radial circlet approximately 38% of aboral cup height. Radial facets angustary, declivate, occupy approximately 33% of distal radial plate width; radial facet topography not known. Three anal plates in cup; radianal heptagonal, in sutural


contact proximally with infrabasal plates, directly below C radial plate; anal X large, above and to left of radianal, wider than high, laterally adjacent to C and D radial plates; right tube plate partially in aboral cup. Tegmen and anal structure unknown. Arms branch isotomously two times. First primibrachial as


high as wide, sixth primibrachial axillary, primaxil as wide as high. First secundibrachials in lateral contact medially, seventh or higher secundibrachial axillary; first tertibrachials not in lateral contact medially. All brachials rectangular uniserial, aborally convex, nonaxillary brachial dimensions variable but wider than high; convex. Column circular, holomeric, and heteromorphic; nodals


alternative with one internodal. Proxistele widest immediately below aboral cup, tapers distally to mesistele where column diameter is constant as known.


Etymology.—Villasi, in recognition of Enrique Villas, who contributed much to understanding Ordovician stratigraphy of the Iberian Chains.


Measurements.—Holotype: crown height, 16.9*; aboral cup height, 4.3; aboral cup distal width, 4.3*; aboral cup proximal width, 2.4; column height, 14.2 (measurements in millimeters, *incomplete or specimen crushed).


Remarks.—Picassocrinus villasi is a relatively small crinoid, but most aspects of its morphology are well preserved. As dis- cussed in the following, Picassocrinus has a very unusual CD interray; its column is also unusual. The proximal columnal is the widest columnal, the proxistele columnals become pro- gressively narrow distally, and mesistele columnals appear to be basically uniform in width (not including the differences between nodals and internodals). This morphology suggests that the proxistele and mesistele were distinct column segments separated by a generating columnal (Wulff and Ausich, 1989); however, clear evidence of a generating columnal is absent. As argued byWulff and Ausich (1989), it seems improbable that a crinoid would insert a wide columnal under that aboral cup only to have it reduce in size through ontogeny. If narrowing the width through ontogeny is not true, two possibilities remain to


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