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Ausich et al.—Disparid and hybocrinid crinoids from the Brechin Lagerstätte


861


zoidal; E inferradial trapezoidal, occupying ~20% of radial- basal articulation; E superradial occupying ~55% of the distal aboral cup margin. Distal facet of A and D radials supporting a lateral arm. Other aspects of radial and anal plates unknown, including verification that B arm is present. Four arms presumably present. Arms slender with aborally


convex brachials; brachials rectilinear uniserial or cuneate uniserial, higher than wide, yielding arms with zig-zag appear- ance. Main axil with nonaxillary brachials; divisions isotomous. E-ray arm unbranched; E-ray brachials higher than wide, rectilinear uniserial (Fig. 8.4). Main axils weakly developed; two axillaries present, each separated by one nonaxillary brachial. Lateral arms apparently with normal bilateral heterotomy. Primaxil arm bifurcating at least through epsilon axillary with two (proximally) or four (distally) nonaxillary brachials before each bifurcation; three divisions preserved on betaaxil arm with three and five nonaxillary brachials before each bifurcation; beta ramules not visible. Other details of arms unknown. Column homeomorphic; proxistele with very thin columnals


with wavy or straight sutures. Beneath aboral cup proxistele columnals approximately six times wider than high and wedge shaped.Mesistele and dististele columnals three times wider than high.Holdfast an irregular, subovatemass of stereomcemented to crinoid pluricolumnal. Holdfast classified as simple, discoidal, cemented type (Fig. 8.4).


Materials.—CMCIP 366334 (3 specimens);UMMP74664 and 74665.1.


Measurements.—UMMP 74664: CrH, 30.0*; ACH, 5.0; proxi- mal, ACW, 7.0; CoH, 24.0. UMMP 74665.1: CrH, 30.0; ACH, 4.5; proximal ACW, 5.1; CoH, 19.5*.


Remarks.—See remarks under genus.


Cremacrinus inaequalis (Billings, 1859) Figure 9.4, 9.5


Figure 7. Anomalocrinus astrictus n. sp., holotype, C-ray lateral view of partial crown (note large brachials in the C-ray arm); calyx sculpture is smooth and pitting on calyx plates on this specimen is a preservational artifact (compare to Figure 5.4), UMMP 74661. Scale bar=5.0mm.


Description.—Crown without perfect bilateral symmetry, probably pendant on column. Aboral cup small, adorally-aborally com- pressed; plate sculpture coarsely pitted; sutures impressed (Fig. 8.5). Basal plates four, all in column concavity, all part of distal


margin of basal circlet. Radial circlet almost twice as wide as high, wider proximally, tapering distally (Fig. 8.5); trapezoidal and slightly indented along line between proximal edge ofAand D radial facets; ligament pit divided. A and D radial plates occupying majority of radial circlet. E-ray inferradial and superradial with narrow sutural contact; E superradial trape-


1859 Heterocrinus inaequalis Billings, p. 51, pl. 4, fig. 7a. 1868 Heterocrinus inaequalis; Shumard, p. 377. 1868 Heterocrinus inaequalis; Bigsby, p. 20. 1886 Cremacrinus inaequalis; Ulrich, p. 113. 1889 Calceocrinus inaequalis; Miller, p. 230. 1889 Castocrinus inaequalis; Ringueberg, p. 395, pl. 10, fig. 5. 1915 Cremacrinus inaequalis; Bassler, p. 289. 1943 Cremacrinus inaequalis; Bassler and Moodey, p. 376. 1946 Cremacrinus inaequalis; Wilson, p. 35, pl. 5, fig. 4. 1962 Cremacrinus inaequalis; Moore, p. 21. 1973 Cremacrinus inaequalis; Webster, p. 86. 1982 Cremacrinus inaequalis; Brower, p. 93. 1988 Cremacrinus inaequalis; Webster, p. 60.


Holotype.—GSC 1444.


Diagnosis.—Cremacrinus with smooth aboral cup sculpture; aboral cup wider than high; aboral cup plates not swollen; axil arms two; axillaries not swollen; arms slender; brachials recti- linear uniserial, wider than high; E-ray arm unbranched.


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