558


Journal of Paleontology 92(4):546–567


Table 4. Comparison between some species of Agathoxylon/Araucarioxylon from Gondwana sharing the characters: presence of circular pits (some flattened) and alternate (opposite) in the radial walls of the tracheids. (1/2: rays uniseriate partially biseriate; in parentheses=character with occasional condition).


RAYS CHARACTERS/SPECIES


A. vesturense Agashe and Prasad A. zaranense Agashe and Prasad A. bengalense (Holden) Maheshwari A. meyenii Agashe and Gowda A. allanii (Kräusel) Maheshwari A. allanii (Kräusel) Maheshwari


A. allanii (Kräusel) Gnaedinger and herbst A. allanii (Kräusel) Maheshwari A. petriellae Zamuner


A. malaimbandense (Marguerier) Gnaedinger and Herbst Araucarioxylon sp. McLoughlin. 1992 A. kumarpurensis Bajpai and Singh A. kumarpurensis Bajpai and Singh


A. kumarpurensis (Bajpai and Singh) Crisafulli and Herbst A. kumarpurensis (Bajpai and Singh) Crisafulli and Herbst A. lamaibandianus Crisafulli and Herbst


Agathoxylon dallonii (Boureau) Crisafulli and Herbst


D.(A.) dallonii (Boureau) Duperon-Laudoueneix and Lejal-Nicol


A. liguanensis Torres and Philippe A. liguanensis Torres and Philippe


A. termieri (Attims) Gnaedinger and Herbst A. termieri (Attims) Gnaedinger and Herbst A. termieri (Attims) Gnaedinger and Herbst A. termieri (Attims) Gnaedinger and Herbst


A. wagadensis Ra, Prasad, Prakash, Singhi, Garg, Gupta, and Pandey


A. sriperumbudurensis Kumarasamy A. protoaraucana Brea


A. protoaraucana (Brea) Gnaedinger and Herbst


A. ohzuanum (Nishida, Ossawa, Nishida, and Rancusi) Gnaedinger and Herbst


A. quiriaquinaense (Nishida) Gnaedinger and Herbst A. chilensis Nishida


RADIAL WALLS PITS


CROSS- FIELD WIDTH HEIGHT


1–41–9, 13 1 (2) 1–26 1–41–8 1–3 (4)


1 (2) 1–21 2–71 1–20 1–41–11


1–2 (3) groups 1–6, 8 1 1–2(3–4)


2–6, 12 1


1 (1/2) 1–18 (–26) 1–12 1–27


1–22–41 3–9(–12) 1–22–41 3–9 2–3(1–4)


9–20


1–21–6 1–2(3)


2 (1.3–4)


1 (1/2) 1–21 1 (1/2) 1–23


1–21–22–34 2–8


1 (1/2) 1–19 1 (2)


1–34–81 1–12 4–71 3–9


2 (1.3)


1–21–41 3–23 2–41 6–17


1–21–2 (3) 1(1/2) 1–12 (–39) 1–21–31 1–30 (56)


1–2.3 1–2.3 1–2.3 1–2.3


1 (2)


2–10 3–10 2–10


1(1/2) 2–8(–20) 1(1/2) 2–6(–14) 1(1/2) 2–14 (–20)


2–8,10 1(1/2) 2–12 (–20)


1–22–51 2–26 3–61 1–15


1–23–91 2–9(–11) Up to 8 1(1/2) 2–12 (–19)


1 (2) 1–22–41 1–11 1–21–4.5 1


Remarks.—The comparisons made with the species of Agathoxylon described for Gondwana justify assignment of the La Matilde Formation specimens to a new species. In Table 5, comparisons made between species of Agathoxylon, Dadoxylon, and Araucarioxylon of Gondwana show that they share the following diagnostic characters: presence of circular, hexagonal, and flattened radial pitting. The newspecies is distinguished from the other Gondwanan


species by the arrangement of the tracheid pits, the number of pits in the cross-field, and by the height of the rays (Table 5). The most closely comparable species is Agathoxylon semibiseriatum


1–21–51 1–10 2–15


ORIGIN/REFERENCE India


Agashe and Prasad, 1989 India


Agashe and Prasad, 1989 India


Holden, 1917; Maheshwari, 1972 India


Agashe and Gowda, 1978


Antarctica, Kräusel, 1962


Antarctica Maheshwari, 1972


Uruguay Crisafulli et al., 2009


Argentina, Crisafulli et al., 2000


Uruguay Zamuner, 1996 Africa


Marguerier, 1976


Australia McLoughlin, 1992


India, Bajpai and Singh, 1986


Africa, Roberts et al., 1997


Argentina Crisafulli and Herbst, 2008


Uruguay Crisafulli et al., 2009


Argentina Crisafulli and Herbst, 2011


Argentina Crisafulli and Herbst, 2011


Africa Boureau, 1948; Duperon-Laudoueneix and Lejal-Nicol, 1981


Chile Torres and Philippe, 2002


1–21–3,4–6 1(1/2) 1–30 (46–56) Argentina Gnaedinger et al., 2015


Africa Attims, 1965


Africa Giraud and Hankel, 1985


Neuquén, Argentina Gnaedinger, 2006


Santa Cruz, Argentina (this work)


India Rai et al., 2016 India


Kumarasamy, 2016


Argentina Brea, 1997


Argentina Gnaedinger and Herbst, 2009 Chile


Nishida et al., 1992 Chile


Nishida, 1984 Chile


Nishida, 1970


Lower Permian Lower Permian Upper Triassic Upper Triassic


Jurassic Jurassic– Cretaceous?


Lower Jurassic


Lower Jurassic


Lower Jurassic


Lower Jurassic


Lower Jurassic


Middle Jurassic


Middle Jurassic


Early Cretaceous Middle Triassic Lower Jurassic Upper


Cretaceous Cenozoic


Cenozoic


Cenozoic, Miocene


(Pant and Singh, 1987) Leiva Verón and Crisafulli in Leiva Verón et al., 2012 inmost of the anatomical characters (Table 5), but there are differences in the radial pitting and rays.


Qualitative and quantitative analysis of anatomical characters


Tables 1 and 2 compare the four species of Agathoxylon iden- tified from the La Matilde Formation, Santa Cruz, Argentina. The anatomical characters must be taken into account for the


AGE


Permian Permian Permian Permian Permian Permian


Lower Permian Permian


Lower Permian


Upper Permian Lower Triassic Upper Permian


Upper Permian


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