664


Journal of Paleontology 92(4):661–680


Remarks.—The suprafamilial classification follows Liebau (2005); for family and other infrafamilial taxa, classification fol- lows Hou et al. (2002), who proposed the family Quad- racyprididae, subfamily Quadracypridinae, encompassing the genera Quadracypris, Nanxiongium, Harbinia,and Sinocypris. The genus Harbinia is widely discussed byDoCarmo et al. (2008) due to its similarity to Pattersoncypris Bate, 1972.The type species Pattersoncypris micropapillosa Bate, 1972 is considered a junior synonym of Harbinia Tsao, 1959. The subspecies described by Krömmelbein and Weber (1971) belonging to Hourcqia Krömmelbein, 1965b (i.e., Hourcqia angulata angulata Kröm- melbein and Weber, 1971; Hourcqia angulata salitrensis Krömmelbein and Weber, 1971; Hourcqia angulata sinuata Krömmelbein and Weber, 1971; and Hourcqia angulata symmetrica Krömmelbein and Weber, 1971) were reassigned to Harbinia and elevated to the level of species. A revision of Hourcqia Krömmelbein, 1965b, Pattersoncypris Bate, 1972, and Harbinia Tsao, 1959 by Poropat and Colin (2012a) led to revali- dation of the genus Pattersoncypris, encompassing the species Harbinia micropapillosa, H. salitrensis,and H. sinuata.Theyalso proposed the new genus Kroemmelbeincypris Poropat and Colin, 2012a, including in it the species Harbinia angulata and H. symmetrica. That proposal was based on the inclined posterior margin, which would differentiate those species from Harbinia. Several characteristics described for Kroemmelbeincypris (e.g., valve overlap, outline, and ornamentation pattern) are also present in Harbinia. Tomé et al., (2014) invalidated the genus Kroemmelbeincypris due to the small variation associated with polymorphism present in some species of Harbinia, and accepted the validity of Pattersoncypris.


Harbinia aff.Harbinia angulata (Krömmelbein and Weber, 1971) Figure 3.1–3.3


1971? Hourcqia angulata angulata Krömmelbein and Weber, p. 81, pl. 6, figs. 23–26.


2002? Pattersoncypris angulata angulata (Krömmelbein and Weber); Coimbra et al., p. 691, fig. 4.29.


2008? Harbinia angulata (Krömmelbein and Weber); Do Carmo et al., p. 795, fig. 6.11.


2012b? Kroemmelbeincypris angulata (Krömmelbein and Weber); Poropat and Colin, p. 709, fig. 4.5.


2014? Pattersoncypris angulata (Krömmelbein and Weber); Tomé et al., p. 165, fig. 10G–I.


Holotype.—A carapace (BfB, type Nr. 7795) from the Post-Bahia Series, Riachuelo layers, Alagoas State, Brazil (Krömmelbein and Weber, 1971).


Occurrence.—Brazil: Sergipe-Alagoas Basin, Riachuelo Forma- tion, upper Aptian (Krömmelbein and Weber, 1971); Araripe Basin, Santana Formation, Romualdo Member, Aptian (Poropat and Colin, 2012b); Cedro Basin, deposits correlated to the Crato Formation, Aptian (Tomé, 2007); Jatobá Basin, Serra Negra, in deposits corre- lated to the Crato Formation, upper Aptian (Tomé et al., 2014); São Francisco Basin, Quiricó Formation, Lower Cretaceous, Aptian.


Materials.—Four intervals from São José creek, São José Farm, Presidente Olegário, Minas Gerais State, southeastern Brazil:


MP-2882, two carapaces; MP-2883, 40 carapaces; MP-2889, 35 carapaces; MP-2922, six carapaces.


Remarks.—This species is easily identified by the marked posterior cardinal angle that coincides with the greatest length of the carapace, forming also a posterior lump in some instars (Tomé et al., 2014). The recovered individuals are poorly preserved, with evidence of dissolution.


Harbinia symmetrica (Krömmelbein and Weber, 1971) Figure 3.4–3.6


1971 Hourcqia angulata symmetrica Krömmelbein and Weber, p. 81, pl. 6, fig. 25.


1990 Cultella sp. 1 Dépêche et al., p. 308, pl. 2, fig. 2.


1990 Pattersoncypris cf. angulata angulata (Krömmelbein and Weber); Musacchio, p. 564, pl. 1, fig. 4.


1990 Hourcqia angulata symmetrica Krömmelbein and Weber; Silva-Telles and Viana, p. 325, pls. 1, 3, fig. 8.


1999 Pattersoncypris angulata symmetrica (Krömmelbein and Weber, 1971); Bate, p. 289.


2002 Pattersoncypris angulata symmetrica (Krömmelbein and Weber); Coimbra et al., p. 691, Fig. 4.30.


2004a Harbinia symmetrica (Krömmelbein and Weber); Do Carmo et al., p. 144, fig. 4.1.


2004a Harbinia sp. 1 Do Carmo et al., p. 144, fig. 4.2.


2006 Harbinia aff. Harbinia symmetrica (Krömmelbein and Weber); Ramos et al., p. 344, fig. 4M–P.


2008 Harbinia symmetrica (Krömmelbein and Weber); Do Carmo et al., p. 795, fig. 6.9.


2012b Kroemmelbeincypris symmetrica (Krömmelbein and Weber); Poropat and Colin, p. 709, fig. 4.4.


Holotype.—A carapace (BfB, type Nr. 7797), from the Codó layers,Maranhão State, Brazil (Krömmelbein and Weber, 1971).


Occurrence.—Brazil: Araripe Basin, Rio da Batateira Forma- tion, and Santana Formation, Crato, Ipubi, and Romualdo members, Aptian (Coimbra et al., 2002), Harbinia spp. 201– 218 Zone, coded as NRT-O11 (Do Carmo et al., 2008), Alagoas Stage (Schaller, 1968; Moura, 1987), upper Aptian–lower Albian (Antonietto et al., 2012); Parnaíba Basin, Codó Forma- tion (Krömmelbein and Weber, 1971; Ramos et al., 2006); São Francisco Basin, Quiricó Formation, Lower Cretaceous, Aptian. Africa: Gabon Basin, Gamba Formation; Congo and Cabinda basins, Chela Formation (Grosdidier et al., 1996; Bate, 1999).


Materials.—Four levels from Quiricó creek, São José Farm, Presidente Olegário, Minas Gerais State, southeastern Brazil: MP-2879, five carapaces; MP-2882, two carapaces; MP-2883, 50 carapaces; MP-2885, two carapaces.


Remarks.—Poropat and Colin (2012b) included this species in the genus Kroemmelbeincypris based on its outline. However, Tomé et al. (2014) invalidated this genus, justifying that these differences resulted from environmentally induced polymorphism.


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