1158
Journal of Paleontology 91(6):1148–1165 Cardites feruglioi can be distinguished from C. partschii
(Goldfuss, 1840) (Miocene from Europe; Sieber, 1954, pl. 1, fig. 3) by having a larger shell with subtriangular outlines, and
narrower radial ribs that become better developed on the ventral and posterior regions of the valves and are separated by wider intercostal spaces.
Genus Kalelia new genus
Type species.—Cardita multicostata Lamarck, 1806. Late Paleocene, Paris Basin, France.
Included species.—K. burmeisteri (Böhm, 1903),K. pectuncularis (Lamarck, 1806).
Diagnosis.—Carditid with large shells for the family, thick and subcircular. Posterior margin convex, large umbones and posterior area defined by an umbonal carina and a convexity change. Lunule large, vertically oriented. Hinge high, with wide ventral development. Right anterior tooth long, inclined back- wards; right middle tooth thick, strongly curved, with broad base. Left anterior tooth triangular, very broad, inclined back- wards, with very convex posterior edge. External sculpture of tripartited radial ribs, 25–30 wide, most prominent on anterior, intercostals spaces confined to a narrow groove.
Etymology.—The name honors Kal-El, Kryptonian name of Superman, the first superhero in comic books. This genus is one of the oldest and largest alticostate carditids.
Remarks.—Kalelia n. gen. is the largest member of all alticos- tate carditids, with shells reaching up to 60 mm in height. It is characterized by the development of a tall hinge plate, promi- nent teeth, a large lunule, and the presence of low radial ribs; subrectangular in cross section. These characters are shared with planicostate carditids, but the development of tripartite radial ribs places Kalelia within alticostates carditids. Rutsch (1936) and McClure (2009) already mentioned the possibility that ‘Venericardia’ pectuncularis convergently developed some planicostate characters. We proposed including in Kalelia n. gen., apart from
the type species ‘Cardita’ multicostata (Lamarck, 1806) (Cossmann and Pissarro, 1904, pl. 31, fig. 97–3), the late Paleocene species ‘C.’ burmeisteri (Böhm, 1903) (Roca and Salamanca formations, early Danian, Argentina) (Fig. 5.1–5.11) and ‘C.’ pectuncularis (Lamarck, 1806) (Cossmann and Pissarro, 1904, pl. 31, fig. 97–2) (Fig. 6.1, 6.2), from the Sables de Bracheux, France. These species have large shells with strong hinge teeth and the particular external sculpture described above. The poorly known species ‘V.’ marylandica (Clark and Martin, 1901) (Clark and Martin, 1901, pl. 40, fig. 7; Nanjemoy
Formation, early Eocene, USA) is possibly a North American representative of this genus. Kalelia n. gen. differs from Venericardia Lamarck, 1801
(La Perna et al., 2017, fig. 3) in having larger shells with more nearly circular outlines, a convex posterior margin instead of a truncated one, a larger lunule, a higher hinge plate, a triangular right anterior tooth, and fewer radial ribs that are separated by intercostal spaces defined by narrow grooves. Kalelia n. gen. can be distinguished from Arcturellina
Chavan, 1951 (type species C. asperula Deshayes, 1825, Eocene, Paris Basin) (Cossmann and Pissarro, 1904, pl. 22, fig. 97-22) by having larger shells, convex posterior margin, higher hinge plate, long or straight right anterior tooth, triangular left anterior tooth, and fewer radial ribs covered with subcircular nodes instead of the subrectangular ones that characterize Arcturellina. Kalelia n. gen. can be distinguished from Claibornicardia
Heaslip, 1968 (Heaslip, 1968, pl. 22, figs. 8–9, pl. 23, figs. 1–2) by having a rounded posterior margin, convex umbones, a well- developed posterior area, the presence of a lunule, the higher number of radial ribs with subcircular nodes, and narrower intercostals spaces separating the ribs.
Stenzel and Krause, 1957 (Heaslip, 1968, pl. 26, figs. 6–11) because Kalelia has larger and more nearly circular shells, a higher and more robust hinge, and fewer, lower, and wide radial ribs. Kalelia n. gen. can be separated from Rotundicardia
Kalelia burmeisteri (Böhm, 1903) n. comb. Figure 5.1–5.7
1902 Cardita morganiana Rathbun; Burckhardt, p. 216, figs. 1,7.
1903 Cardita burmeisteri Böhm, p. 72. 1903 Cardita Burckhardti Ihering, p. 205. 1907 Venericardia burmeisteri (Böhm); Ihering, p. 23. 1938b Venericardia burmeisteri (Böhm), ‘Forma B’; Fossa- Mancini, p. 263.
1967 Venericardia paleopatagonica var. australis Masiuk, p. 243, pl. 5, fig. 2.
Type specimens.—Holotype MLP 15294a and 15294f. Two internal molds from General Roca (Río Negro Province).
Diagnosis.—Shell medium-sized, posterior margin slightly convex, dorsal margin sloping from beaks to posterior one, posterior area sculptured by three or four higher and wide radial ribs, continued by three lower and narrower other ones. Umbones subcentrally placed. External sculpture of 25 wide and low radial ribs.
Occurrence.—General Roca area (early Danian, Roca Forma- tion, Río Negro Province) and Puesto Álvarez (early Danian, Salamanca Formation, Chubut Province).
Figure 5. Kalelia burmeisteri (Böhm, 1903) n. comb. (1–7): (1) external view of left valve, CPBA 8338, from Puesto Álvarez; (2) external view of left valve with shell removed, CPBA 8338, from Puesto Álvarez; (3) external view of right valve, CPBA 8338, from Puesto Álvarez; (4) internal view of right valve, MACN-Pi 5357, from General Roca; (5) left view of internal mold, CPBA 8338, from Puesto Álvarez; (6) left view of internal mold, holotype MLP 15294f, from General Roca; (7) left view of internal mold, holotype MLP 15294a, from General Roca. Kalelia multicostata (Lamarck, 1806) n. comb. (8–11): (8) internal view of left valve, holotype MNHN A07711, from Abbecourt, France; (9) external view of left valve, holotype MNHN A07711, from Abbecourt, France; (10) internal view of right valve, MNHN F07559, from Noailles, France; (11) external view of right valve, MNHN F07559, from Noailles, France. Scale bar represents 10mm.
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