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1202


Journal of Paleontology 91(6):1199–1219


Figure 2. History of different conodont zonal schemes documented by different authors for the Wuchiapingian Stage.


in the Penglaitan and Fengshan sections, Guangxi, and in the Xiaoyuanchong Section, Hunan (Mei et al., 1994b). Zhang et al. (2008) recognized the Clarkina postbitteri postbitteri Zone in the Maoershan Section, Hubei. However, some specimens were


illustrated thatmight have serrations (e.g.,Zhang et al., 2008, pl. 2, fig. 4), which is a character belonging to Jinogondolella.Other


specimens may be Clarkina leveni (e.g., Zhang et al., 2008, pl. 3, fig. 10) in samples from the C. transcaucasica Zone of Zhang et al. (2008). Thus, almost all specimens illustrated in Zhang et al. (2008) cannot be identified as C. postbitteri postbitteri and this zone is doubtful in the Maoershan Section. Some C. postbitteri postbitteri specimens were identified as C. dukouensis by Wang (2000), who did not recognize this zone in the Penglaitan Section (see Systematic paleontology for details). Shen and Zhang (2008) recognized this zone in the uppermost part of Douling Formation in Chenzhou,Hunan. Thus, this zone has only been recognized in a few regions of South China.


Clarkina dukouensis Zone.—This zone is defined by the first


occurrence of Clarkina dukouensis at the base and by the first occurrence of C. asymmetrica at the top, and was first established in the Dukou and Nanjiang sections, Sichuan (Mei et al., 1994a). The Clarkina dukouensis Zone is the first Wuchiapingian conodont zone in most regions of South China, due to the global transgression following the Guadalupian- Lopingian lowstand interval. However, in some sections, only more advanced specimens of C. dukouensis are recognized in the basal part of the Wuchiaping Formation.


Clarkina asymmetrica Zone.—This zone is defined by the first occurrence of Clarkina asymmetrica at the base and by the first


occurrence of C. leveni at the top, and was first established in the Dukou and Nanjiang sections, Sichuan (Mei et al., 1994a).


Clarkina leveni Zone.—This zone is defined by the first occur- rence of Clarkina leveni at the base and by the first occurrence of C. guangyuanensis at the top, and was first established in Archura, Transcaucasia (Kozur, 1975). It has been recognized in northwest Iran (Henderson et al., 2008; Shen and Mei, 2010). The Clarkina leveni Zone cannot be recognized in a few sections, but the reason is not clear. For example, it was not recognized in the Penglaitan Section, but it was in the Tieqiao Section, which is just 10 km west of the Penglaitan Section.


Clarkina guangyuanensis Zone.—This zone is defined by the first occurrence of Clarkina guangyuanensis at the base and by


the first occurrence of C. transcaucasica at the top, and was first established in the Dukou and Nanjiang sections, Sichuan (Mei et al., 1994a). The marker species, C. liangshanensis, began to occur in this zone. However, the range of C. liangshanensis extends to the middle of the C. orientalis Zone.


Clarkina transcaucasica Zone.—This zone is defined by the first


occurrence of Clarkina transcaucasica at the base and by the first occurrence of C. orientalis at the top, and was first estab- lished in the Dukou and Nanjiang sections, Sichuan (Mei et al., 1994a). Abundant C. liangshanensis can coexist with Clarkina transcaucasica in this zone.


Clarkina orientalis Zone.—This zone is defined by the first occurrence of Clarkina orientalis at the base and by the first occurrence of C. wangi at the top. Mei et al. (1994a) established


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