1206


Journal of Paleontology 91(6):1199–1219


anterior 1/3 of the element; some specimens have the widest point in the middle portion, narrowing gradually both posteriorly and anteriorly. Posterior end of platform is bluntly rounded, but more upturned. Cusp is erect, terminal, obviously higher and larger than the posterior and middle denticles, and sometimes fused with the posteriormost denticle. It is slightly reclined in juveniles. Tight and numerous denticles are almost equal in height except for those in the anterior in most individuals, or increasing gradually in height anteriorly in juveniles, and the middle denticles on the carina are more fused in gerontic specimens, but less in junior ones. Furrows are moderately wide and smooth. Serrations are in the anterior 1/3 or 1/4 in most specimens, especially in gerontic forms, but not obvious in juveniles and some adults.


Remarks.—Jinogondolella granti is the youngest species of Jinogondolella in South China. It can be differentiated from J. xuanhanensis by its bluntly rounded and upturned posterior end, more fused middle carina, and nearly parallel lateral margins, but J. xuanhanensis has a flat, concave and slightly deflected posterior end of platform. Jinogondolella granti is similar to J. shannoni, however the latter has more discrete denticles than the former in the middle part, and has the widest point in the posterior portion narrowing gradually anteriorly in almost individuals. Wang (2000) identified one specimen as Clarkina postbitteri


in Bed 4 at the Penglaitan Section (in the J. granti Zone), but itwas determined to be sample contamination (Wang, 2000, p. 6).Many specimens were identified by Wang (2000) as J. altudaensis, J. shannoni,and J. prexuanhanensis, which coexisted with J. granti at the Penglaitan Section. However, the characteristics of almost all of those individuals coincide with J. granti according to the diagnosis, therefore,theyare allassignedto J. granti herein. Wang (2000) established a new species, Mesogondolella laibinensis, in Bed 4, based on the major character of lacking serration. However, despite not having obvious serration, these specimens still have a slender, symmetrical and long platform, bluntly rounded and upturned posterior end, more fused middle carina, and nearly parallel lateral margins, which also coincide with those of Jinogondolella granti (see emended diagnosis and Fig. 3.5, 3.6, 3.11, 3.12). Thus, Mesogondolella laibinensis is considered to be a synonym of Jinogondolella granti. Almost all


specimens identified as J. granti by Zhang et al. (2008) still have a flat and slightly deflected posterior end of the platform, which characterizes J. xuanhanensis; they may be from the samples with transitional J. xuanhanensis to J. granti.


Genus Clarkina Kozur, 1989


Type species.—Gondolella leveni Kozur, Mostler, and Pjatakova in Kozur, 1975 from Achura, Azerbaijan.


Remarks.—Clarkina has been discussed in detail by Yuan et al. (2014a). In addition to the oral surface, Kozur (1989) also illustrated the aboral surface, which he considered to be different from Triassic Gondolellidae genera, but similar to Permian Gondolellidae genera, with the distinct difference primarily in the oral surface among Gondolellidae genera (e.g., free blade). Thus, the aboral surface difference may be a diagnostic character at genus level, but not for the Permian Gondolellidae genera.


The aboral surface is not illustrated because this paper only focuses on the difference amongWuchiapingianClarkina species.


Clarkina postbitteri hongshuiensis Henderson, Mei, and Wardlaw, 2002 Figure 3.19–3.34


?1998a Clarkina postbitteri; Mei et al., pl. 5, figs. 1, 7. 2000 2002


Clarkina postbitteri; Wang, pl. 1, figs. 10, 11, 14–18.


2006a Clarkina postbitteri hongshuiensis; Jin et al., figs. 7.1, 7.6–7.10, 7.13–7.16.


non 2010 figs. 9, 12.


Clarkina postbitteri hongshuiensis Henderson et al., p. 730, pl. 1, figs. 1–11, pl. 2, figs. 9, 10, 12, 13.


?2006 Clarkina postbitteri hongshuiensis; Sun and Xia, pl. 1, figs. 7–9.


Clarkina hongshuiensis; Wardlaw and Nestell, pl. 15,


Holotype.—NIGP134579, from the Maokou Formation, Guangxi Province, China (Henderson et al., 2002, pl. 1, fig. 5).


Original diagnosis.—A subspecies of Clarkina postbitteri that exhibits a wide range of morphotypes, but always has smooth anterior margins. Almost all mature specimens have a narrow brim and high, fused anterior denticles forming a blade. The diagnostic aspect of this taxon (and other taxa) appears in larger adult forms; small juvenile specimens cannot be used for species discrimination. Some morphotypes show rounded posterior terminations whereas others are more blunt or square with rounded corners. In some specimens, the platform narrows abruptly anteriorly, but in many the platform narrows somewhat more gradually. The number of posterior and middle denticles is rather variable and in some they are mostly discrete, but in others most of the middle denticles become closely spaced to fused. In most specimens, there is a gap between the posterior denticle and the cusp. Some specimens show gaps between the posterior denticles, but in many the posterior denticles are very tightly spaced to fused (Henderson et al., 2002, p. 730).


Emended diagnosis.—A subspecies of Clarkina postbitteri characterized by a P1 element with a slender, symmetrical and slight arched platform whose lateral margins are nearly parallel in the posterior in most individuals, then narrowing gradually in the anterior 1/3 or 1/4 of the element, except for a few specimens in which their platform narrows abruptly anteriorly. Few specimens have the widest point in the posterior end narrowing gradually anteriorly. Posterior end of platform is bluntly rounded, but some larger specimens have a squarely rounded posterior end. Cusp is erect, terminal, and higher and larger than the posterior denticles in most individuals, but is nearly same height as the posterior denticles in gerontic specimens,which usually have a narrowbrim in the posterior end. Posterior and middle denticles are closely spaced, but unfused, whereas in juveniles they are more discrete and form gaps. Posterior denticles are about equal in height, then increase gradually and become more fused anteriorly, until they form a free blade in the anterior. The gap between the cusp and the posteriormost denticle is not obvious. Furrows are narrow and smooth in most individuals.


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