1194
Journal of Paleontology 91(6):1178–1198
is of middle Fortunian age, and is defined based on the first appearance of Treptichnus pedum and the earliest bilobate trace Cruziana isp. Ichnozone 3 is late Fortunian–Cambrian Age 2, and is marked by sudden change in abundance and complexity of trace fossils and the first appearance of Cruziana pro- blematica. Ichnozone 4 is Cambrian Ages 2–3, and is char- acterized by the first appearance of Psammichnites gigas, Rusophycus avalonensis, and Didymaulichnus miettensis.An integration of trace fossils with small shelly fossils and evalua- tion suggest that the Ediacaran-Cambrian boundary should be placed at the base of the Soltanieh Formation or within the Lower Dolomite Member.
Acknowledgments
Financial support for this study was provided by Natural Sci- ences and Engineering Research Council (NSERC) Discovery Grants 311727–08/15 and 311726–13 awarded to Mángano and Buatois, respectively, as well as a University of Saskatchewan President Fund to Mángano. A. CiabeGhodsi is thanked for introducing us to the area and providing valuable feedback. A. EbrahimNejad and H. Behmaram helped the first author during fieldwork. Reviewers S. Jensen and M. Laflamme and Associate Editor L. Tapanila provided very useful comments that helped to improve the manuscript.
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