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Journal of Paleontology 91(6):1178–1198
favors inclusion in T. pollardi. Treptichnus pollardi has been typically, though not exclusively (e.g., Geyer and Uchman, 1995), recorded from freshwater environments (Buatois and Mángano, 1993a, 1993b; Fregenal-Martinez et al., 1995; Metz, 1995; Buatois et al., 2000).
Occurrence.—Upper Shale Member.
Treptichnus isp. Figure 5.2
Materials.—Four slabs (P3310.1, P3310.2, P3311.9, P3319.2) containing seven specimens.
Description.—Horizontal, gently curved, branching burrows. The burrows occasionally exhibit a short projection. Width is 1.1–2.6mm. Preserved as positive hyporelief.
Remarks.—Lack of zigzag pattern and regular projection does not allow ichnospecific assignment.
Occurrence.—Upper Shale Member. Previous work
In order to contextualize the new trace-fossil data reported here and to define a biozonation for the Soltanieh Formation, pre- viously published reports of trace fossils and SSFs from this unit are briefly reviewed and critically re-evaluated.
Lower Dolomite Member.—Hamdi et al. (1989) reported phosphatic tubes, including Hyolithellus sp. along with others resembling Rugatotheca sp., fragments of the protoconodont Protohertzina sp., and globomorphs of the Olivooides multisulcatus Qian, 1977 group in the upper part of the Lower Dolomite Member in the Vali-Abad area.
Lower Shale Member.—Hamdi et al. (1989) and CiabeGhodsi (2007) recorded large discoidal algal vesicles assigned to Chuaria sp. from the lower interval of the Lower Shale Member in the Vali-Abad area, and the type section, respectively. Hamdi et al. (1989) compared their material with larger ellipsoidal vesicles of Shouhsienia sp. from the Sinian System in China. CiabeGhodsi (2007) reported Diplocraterion isp., Planolites vulgaris, Skolithos isp., and Treptichnus pedum from the upper interval of the Lower Shale Member. Planolites vulgaris needs re-evaluation because the type specimen of this ichnospecies has been regarded as inorganic (Pemberton and Frey, 1982). Based on the illustration provided, an affinity with Hel- minthoidichnites cannot be disregarded. In addition, specimens attributed to Diplocraterion isp. and Skolithos isp. are based on bedding-plane expressions, so their true morphology cannot be confirmed.
Middle Dolomite Member.—Hamdi et al. (1989) reported the tubular fossil Hyolithellus vladimirovae and protoconodonts of the Protohertzina anabarica Missarzhevsky, 1969 group from the lower interval of the Middle Dolomite Member. Less com- mon elements include the tubular fossils Anabarites trisulcatus
Missarzhevsky in Voronova and Missarzhevsky, 1969; Cam- brotubulus decurvatus Missarzhevsky in Rozanov et al., 1969; siphogonuchitids; Palaeosulcachites sp.; Siphogonuchites sp.; and globomorphs. These authors mentioned that beds near the top of the Middle Dolomite Member contain a similar assem- blage, but with more abundant A. trisulcatus,C. decurvatus, and Siphogonuchites sp., in addition to the primitive mollusks Maikhanella multa and Purella sp., and the tubular fossils Tiksitheca licis Missarzhevsky in Rozanov et al., 1969; Circotheca sp.; and Ladatheca isp.; as well as hyolithids. Tashayoee et al. (2012) recently reported Anabarites latus Val’kov and Sysoev, 1970; A. rectus Vasil’eva in Rudavskaya and Vasil’eva, 1984; A. tripartitus Missarzhevsky in Rozanov et al., 1969; A. trisulcatus; Cambrotublus isp.; Conotheca sub- curvata Yu, 1974; Drepanochites dilatatus Qian and Jiang in Luo et al., 1982; Hyolithellus vladimirovae Missarzhevsky in Rozanov and Missarzhevsky, 1966; Jakutiochrea lenta Val’kov, 1987; Protohertzina anabarica Missarzhevsky, 1969; P. siciformis Missarzhevsky, 1973; P. unguliformis Mis- sarzhevsky, 1973; Siphogonuchites triangularis Qian, 1977; Siphogonuchites sp.; and Yunnanodus dolerus Wang and Jiang in Jiang, 1980 from the Garmab section. CiabeGhodsi (2007) documented the ichnotaxa Bergaueria perata Prantl, 1945; Circulichnus montanus (Vyalov, 1971); and Gordia arcuata from the lower interval of the Middle Dolomite Member at the type section. However, specimens assigned to Bergaueria perata and Circulichnus montanus are unconvincing based on the available material.
Upper Shale Member.—Hamdi et al. (1989) mentioned that at the Vali-Abad section the lower interval of the Upper Shale Member contains abundant and diverse phosphatized mollusks comparable with those found in the upper interval of the Middle Dolomite Member. These authors also reported Anabarites cf. trisulcatus Missarzhevsky, 1969; allathecidae hyoliths; and pelagiellids from the Dalir section. The upper interval in the Vali-Abad section contains specimens of the Latouchella korobkovi (Vostokova, 1962) group of monoplacophorans, including ‘close-coiled’ Yangtzespira sp., ‘lax coiled’ Bemella sp., ‘uncoiled’ Ceratoconus sp., and Obtusoconus sp., which appear approximately 20m from the top of the unit. Other typical elements at this level include Purella tianzhushanensis Yu, 1979 and broad monoplacophorans resembling Proto- wenella sp.; pelagiellids of the Pelagiella lorenzi (Kobayashi, 1939) occur in the top 10 m. CiabeGhodsi (2007) added Lopo- chites latazonalis Qian, 1977; Igorella emeiensis (Yu, 1987); Igorella sp.; Purella squamulosa Qian and Bengtson, 1989; Bemella simplex Yu, 1979; and Lapworthella sp. to this group. CiabeGhodsi (2007) described several trace fossils from the Garmab section, namely Chondrites furcatus Sternberg, 1833; Didymaulichnus miettensis; Diplichnites isp.; Diplocraterion isp; Hormosiroidea isp.?; Monomorphichnus lineatus Crimes et al., 1977; Neonereites uniserialis Seilacher, 1960; Palaeo- phycus alternatus; Paleodictyon croaticum Uchman, 1995b; Plagiogmus arcuatus Glaessner, 1969; Psammichnites gigas; and Protovirgularia dichotoma?M’Coy, 1850. However, many of these ichnotaxa lack diagnostic features and have been excluded from the list of trace fossils used in this study. Struc- tures assigned to Chondrites furcatus do not display the classic
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