O’Dogherty et al.—Jurassic radiolarians from the Eastern Alps


Diagnosis.—Large cryptothoracic tricyrtid with very well defined lumbar stricture and a large abdomen. Outer surface without nodes and covered by hexagonal pore frames. Distal aperture constricted circular without rim.


Occurrence.—Upper Bajocian to lower Bathonian.


Description.—Cryptothoracic subspherical form composed of three distinct segments. Cephalis and thorax partially incased in the abdomen, with a very well defined lumbar stricture. The thorax is truncate-conical and covered by small pores whereas the abdomen is quite large and spherical. The entire surface of the abdomen is covered by a latticed meshwork of polygonal pore frames and a constricted aperture is visible.


Etymology.—The name Nonaginta means ninety, in honor of the prodigious decade in contemporary history of radiolarian research.


Measurements.—(in micrometers; µm) maximum width of shell 112−142, mean 118; maximum length 113−133, mean 123, based on four specimens.


Remarks.—Hemicryptocapsa nonaginta n. sp. was published in open nomenclature in the Mesozoic radiolarian atlas (Baum- gartner et al., 1995b) and since then it has been illustrated in many publications. This species differs from H. yaoi by having larger circular pores in the middle of polygonal areas. It differs from H. carpathica by having a simple circular aperture without a rim.


Hemicryptocapsa yaoi (Kozur, 1984) Figure 9.31−9.37


1984 Praezhamoidellum yaoi Kozur, p. 53, pl. 3, fig. 3a−b. 2003


2004 2006


2006 Williriedellum dierschei; Auer et al., fig. 6.52.


Williriedellum yaoi;O’Dogherty et al., p. 446, fig. 9, figures 6−12. [See for complete synonymy]


2007b Williriedellum dierschei; Gawlick et al., fig. 8.45, 17.32. 2009


Williriedellum dierschei; Suzuki and Gawlick, p. 179, figs. 5.27A–B, 5.28, 6.48A–B.


2010 Williriedellum yaoi; Robin et al., pl. 3, figs. 18, 20.


2012 Hemicryptocapsa yaoi; Goričan et al., pl. 1, fig. 25; pl. 2, fig. 12.


Genus Williriedellum Dumitrica, 1970 Type species.—Williriedellum crystallinum Dumitrica, 1970.


Included species.—CryptamphoreIla crepida O’Dogherty, 1994; Hemicryptocapsa polyhedra Dumitrica, 1970; Hemicryptocapsa prepolyhedra Dumitrica, 1970; Hemicryptocapsa tuberosa Dumitrica, 1970; Sethocapsa yahazuensis Aita, 1987; Tricolocapsa clivosa Aliev, 1967; Tricolocapsa formosa Chiari,


Williriedellum dierschei Suzuki and Gawlick; Suzuki and Gawlick, p. 201, fig. 6.73. [premature name]


Williriedellum dierschei Suzuki and Gawlick in Gawlick et al., p. 311, pl. 4, figs. 1–6.


Emended diagnosis.—The genus Williriedellum originally contained cryptothoracic tricyrtids with an aperture regardless of the external ornamentation of the shell.We restrict the name Williriedellum to species with raised ridges or nodes on the outer surface of the abdomen. Species with a regular distribution of circular pores (each pore may be bounded by a simple polygonal pore frame) are assigned to Hemicryptocapsa. The same differentiation between these two genera was applied by O’Dogherty et al. (2009). The concept of that publication, however, did not allow for written definitions and comments.


Occurrence.—Upper Aalenian to lower Coniacian.


Williriedellum crystallinum Dumitrica, 1970 Figure 9.50−9.51


1970 Williriedellum crystallinum Dumitrica, p. 69, pl. 10, figs. 60 a–c, 62–63.


2009 Williriedellum crystallinum; Suzuki and Gawlick, p. 178, fig. 5.24. [See for complete synonymy]


Williriedellum sp. cf. W. formosum (Chiari, Marcucci, and Prela, 2002) Figure 9.45−9.48


2002 Tricolocapsa formosa Chiari, Marcucci, and Prela, p. 83, pl. 5, figs. 3−8.


2011 Williriedellumformosum; Bandini et al., pl. 5, figs. 27–28. 2012 Williriedellum formosum; Goričan et al., pl. 1, fig. 23.


Remarks.—The external ornamentation of this species is very close toW. formosum but has a rounded aperture whereas in the type material the aperture is covered by a dish-like appendage. Moreover, some of our specimens also have groups of four small pores per frame area like occurring in W. gilkeyi.


Williriedellum yahazuense (Aita, 1987) Figure 9.43−9.44


1987 Sethocapsa yahazuense Aita, p. 73, pl. 2, figs. 8a−9b; pl. 9, figs. 16−17.


1993 Williriedellum sujkowskii Widz and De Wever, p. 88, pl. 1, figs. 7−10.


2005 Williriedellum yahazuense; Šmuc and Goričan, p. 62, pl. 3, fig. 19.


Williriedellum sp. A Figure 10.23−10.24


Remarks.—These specimens show very small and pointed nodes on the outer surface of the abdomen.


49


Marcucci, and Prela, 2002; Williriedellum? gilkeyi Dumitrica, 1970;Williriedellum crystallinum Dumitrica, 1970; Williriedellum nodosum Chiari, Marcucci, and Prela, 2002; Williriedellum peterschmittae Schaaf, 1981; Williriedellum sujkowskii Widz and De Wever, 1993.


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