O’Dogherty et al.—Jurassic radiolarians from the Eastern Alps
Diagnosis.—Tetracyrtid nassellarian having a globose postabdominal segment. Lattice meshwork with polygonal pore frames lacking nodes or tubercles on its surface. Closed antapically or having a discrete number of small pores grouped into a depression (see Sethocapsa uterculus in Baumgartner et al., 1995b, p. 505, pl. 5462, fig. 2). Collar and lumbar strictures slightly recognizable or indistinct externally. Large and globose postabdominal segment is well distinguished from the abdomen by a marked stricture.
Etymology.—From the latin croco (saffron) and capsa (box); feminine gender.
Occurrence.—Bathonian to middle Albian.
Remarks.—O’Dogherty (1994) erected the genus Hiscocapsa for a heterogeneous group of tetracyrtid nassellarians displaying a wide variety of pore frame arrangement. However, many species assigned originally to Hiscocapsa are clearly separated by lacking a distal aperture and bearing no tubercles on the surface. Crococapsa n. gen. differs from Minocapsa Matsuoka,
1991a, by its distinct fourth segment, which is globose and well differentiated from the abdomen by a marked stricture. Crococapa n. gen. also differs from Lantus Yeh, 1987a by having only one postabdominal chamber.
Crococapsa tansinhoki (Hull, 1997) Figure 8.46−8.47
1997 Minocapsa(?) tansinhoki Hull, p. 148, pl. 38, figs. 4, 6.
2006 Minocapsa(?) tansinhoki;O’Dogherty et al., 2006, p. 444, pl. 6, figs. 16, 17. [See for complete synonymy]
Remarks.—O’Dogherty et al. (2006, p. 444) first questioned the placement of this species to Minocapsa because the basal aperture was absent and the phylogentic relationship with early Jurassic representatives was not demonstrated. This species fits well in the general description of the new genus Crococapsa.
Crococapsa sp. aff. C. tansinhoki (Hull, 1997) Figure 9.17−9.18
Remarks.—These forms differ from Hull’s material by having smaller and more numerous pores on the abdomen.
Crococapsa sp. aff. C. truncata (Wu, 2000) Figure 8.48
aff. 2000 Minocapsa truncata Wu, p. 304, pl. 2, figs. 1, 5–6.
Remarks.—This specimen differs from the Tibetan specimens by displaying a proportionally larger last segment.
Crococapsa sp. A Figure 8.49−8.52
Remarks.—These specimens resemble Crococapsa sp. aff. C. truncata (Wu, 2000), but the proximal part of the shell is relatively longer and elongated.
Crococapsa sp. B Figure 8.45
Remarks.—This specimen is closely related to other congeneric forms included in this paper, but differs by the well-marked polygonal pore frames on almost all the shell surface.
Crococapsa sp. C Figure 8.43−8.44
Remarks.—These specimens are very close to Crococapsa tansinhoki but they display more cylindrical and smoother thorax.
Genus Doliocapsa new genus
Type species.—Stichomitra (?) stecki O’Dogherty, Goričan, and Dumitrica in O’Dogherty et al., 2006.
Other species.—Hiscocapsa acuta Hull, 1997; Hiscocapsa minuta Yeh, 2011; Hiscocapsa planata Wu, 2000 (syn. Stichocapsa magnipora); Sethocapsa lugeoni O’Dogherty, Goričan, and Dumitrica in O’Dogherty et al., 2006; Sethocapsa taukhaensis Kemkin and Taketani, 2004; Solenotryma keni Kocher, 1981; Stichocapsa magnipora Chiari, Marcucci, and Prela, 2002; Stichomitra doliolum Aita in Aita and Okada, 1986; and ?Quarticella hunzikeri O’Dogherty, Goričan, and Dumitrica in O’Dogherty et al., 2006.
Figure 8. Scale bars (a, b) measure 50 µm; each radiolarian scanning micrograph is designated by sample and specimen number, followed by the corresponding scale. (1−3) Transhsuum brevicostatum (Ožvoldová), sample BMW-26 (1, 013/b); sample BMW-28 (2, 068/a); sample BMW-34 (3, 002/a). (4−8, 10−18) Transhsuum maxwelli (Pessagno), BMW-13c (4, 083/a; 6, 021/a; 7, 048/a; 18, 024/a); sample BMW-35 (5, 003/a; 13, 027/a; 15, 004/a; 16, 002/a); sample BMW-26 (8, 015/a; 11, 060/a); sample BMW-28 (10, 071/a); sample EW-158 (12, 062/a; 14, 012/a; 17, 061/a). (9) Transhsuum sp. E sensu (Yao), sample BMW-28 (9, 066/a). (19) Eucyrtidiellum pyramis (Aita in Aita and Okada), sample BMW-33 (19, 019/b). (20) Eucyrtidiellum ptyctum (Riedel and Sanfilippo), BMW-13c (20, 139/b). (21−23) Eucyrtidiellum nodosum Wakita, sample BMW-13c (21, 074/b; 22, 005/b; 23, 134/b). (24−25) Eucyrtidiellum unumaense (Yao), sample BMW-28 (24, 017/b); sample BMW-31 (25, 045/b). (26−27) Eucyrtidiellum pustulatum Baumgartner, sample BMW-33 (26, 028/b); sample BMW-13b (27, 060/b). (28) Spinosicapsa basilica (Hull), sample BMW-13c (28, 064/a). (29) Spinosicapsa sp. cf. S. triacantha (Fischli), sample BMW-13c (29, 046/a). (30) Spinosicapsa lata (Yang), sample BMW-33 (30, 018/a). (31) Spinosicapsa spinosa (Ožvoldová), sample BMW-13c (31, 057/a). (32) Yamatoum sp. A, sample BMW-28 (32, 047/b). (33–34) Doliocapsa matsuokai (Yeh), sample BMW-13b (33, 049/b); sample BMW-31 (34, 048/b). (35−37) Doliocapsa planata (Wu), sample BMW-13c (35, 131/a; 37, 089/a); sample BMW-34 (36, 029/a). (38−42) Yamatoum spinosum (Takemura), sample EW-158 (38, 022/b); sample BMW-13b (39, 054/b); sample BMW-26 (40, 073/b; 41, 049/b); sample BMW-28 (42, 051/b). (43–44) Crococapsa sp. C, sample EW-158 (43, 067/b; 44, 070/b). (45) Crococapsa sp. B, sample BMW-35 (45, 032/b). (46–47), Crococapsa tansinhoki (Hull), sample BMW-33 (46, 023/b); sample BMW-35 (47, 022/b). (48) Crococapsa sp. aff. C. truncata (Wu), sample BMW-28 (48, 057/b). (49−52) Crococapsa sp. A, sample BMW-13c (49, 009/b; 50, 073/b; 51, 007/b; 52, 004/b). (53) Belleza decora (Rüst), sample BMW-13c (53, 012/b).
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