O’Dogherty et al.—Jurassic radiolarians from the Eastern Alps
Diagnosis.—Shell with four segments (rarely more), generally thick-walled. Cephalis small, spherical, poreless, without apical horn. Other segments latticed with polygonal pore frames and lacking tubercles or nodes. Last segment somewhat inflated and having an aperture.
Etymology.—From dolium (barrel) and capsa (box); feminine gender.
Occurrence.—Lower Toarcian to Tithonian.
Remarks.—This new genus is differentiated from Praewillir- iedellum Kozur, 1984, by having very distinct strictures and well-differentiated segments.
Doliocapsa matsuokai (Yeh, 2009) Figure 8.33−8.34
1998 Stichocapsa sp. F Arakawa, p. 63, pl. 6, fig. 274. 2005 Stichocapsa sp. Šmuc and Goričan, pl. 3, figs. 22a−b. 2009 Hiscocapsa matsuokai Yeh, p. 67, pl. 21, figs.1,8,20, 22. 2011 Hiscocapsa matsuokai; Yeh, p. 16, pl. 7, figs. 10−13. 2011 Hiscocapsa cf. matsuokai; Yeh, p. 16, pl. 5−7.
Remarks.—The proximal part in our specimens is slightly longer than in the type material. This species differs from all others representatives herein illustrated by having smaller and more numerous polygonal pores, less marked strictures at joints, and longer proximal conical part.
Doliocapsa keni (Kocher, 1981) Figure 7.4
1981 Solenotryma keni Kocher, p. 91, pl. 16, figs. 11−12.
2006 Stichomitra (?) keni (Kocher); O’Dogherty et al., p. 442, pl. 5, fig. 6.
Doliocapsa planata (Wu, 2000) Figure 8.35−8.37
2000 Hiscocapsa planata Wu, p. 303, pl. 1, figs 7–8.
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2002 Stichocapsa magnipora Chiari, Marcucci and Prela, p. 76, pl. 3, figs. 13−17.
Genus Hiscocapsa O’Dogherty, 1994 Type species.—Cyrtocapsa grutterinki Tan, 1927.
Other species.—Sethocapsa aitai Chiari,Marcucci, and Prela, 2002; Sethocapsa kodrai Chiari, Marcucci, and Prela, 2002; Stichocapsa pseudornata Tan, 1927; and Stichocapsa rutteni Tan, 1927.
Emended diagnosis.—Hiscocapsa is emended in order to include only tetracyrtid nassellarians having a globose post- abdominal segment with
tubercles.Alarge distal aperture is also presented and sometimes, when preserved, an appendage. We note that terminally closed species with tubercles (e.g., Setho- capsa kaminogoensis Aita in Aita and Okada, 1986; Sethocapsa spp., pl. 5, fig. 15 in Aita and Okada, 1986; Sethocapsa sp. 8, pl. 2, fig. 32 in Matsuoka, 1998) should belong to a new genus, which is not described herein.
Occurrence.—Upper Bathonian to upper Aptian.
Hiscocapsa kodrai (Chiari, Marcucci, and Prela, 2002) Figure 9.53
2002 Sethocapsa kodrai Chiari, Marcucci and Prela, p. 74, pl. 3, figs. 1−7.
2003 Quarticella ovalis Takemura; Suzuki and Gawlick, p. 199, fig. 5.40.
2005 non Sethocapsa kodrai; Nakae and Komuro, fig. 4.47. 2012 Hiscocapsa kodrai; Djeric et al., pl. 1, fig. 23.
Remarks.—This species is very similar to Quarticella ovalis Takemura, but its aperture is much narrower. The larger postabdominal segment also displays well-differentiated tubercles instead of the typical spiny surface of Quarticella ovalis. Hiscocapsa kodrai might belong to Quarticella, but study of the internal structure is required before this new generic assignation is proposed. In the type material, a flattened base with a constricted aperture is clearly distinct (see Chiari et al., 2002, pl. 3, fig. 3). This flattened base is never present in representatives of Quarticella.
Figure 9. Scale bar measures 50 µm; each radiolarian scanning micrograph is designated by sample and specimen number. (1−6) Hemicryptocapsa buekkensis (Kozur), sample BMW-28 (1, 029; 2, 028); sample BMW-26 (3, 033); sample BMW-31 (4, 032); sample BMW-32 (5, 018); sample BMW-30 (6, 010). (7–8) Zhamoidellum sp. A, sample BMW-32 (7, 051); sample BMW-33 (8, 031). (9) Kilinora? oblongula (Kocher in Baumgartner et al.), sample BMW-32 (9, 005). (10–11) Zhamoidellum sp. aff.Z. ovum Dumitrica, sample BMW-30 (10, 015); sample BMW-32 (11, 086). (12−16) Hemicryptocapsa marcucciae (Cortese), sample BMW-32 (12, 087); sample BMW-26 (13, 074); sample BMW-13c (14, 141); sample BMW-13b (15, 065); sample BMW-30 (16, 043). (17–18) Crococapsa sp. aff. C. tansinhoki (Hull), sample BMW-13c (17, 140; 18, 011). (19) Hemicryptocapsa nonaginta n. sp., BMW-13b (19, 039, paratype). (20−23) Hemicryptocapsa carpathica (Dumitrica), sample BMW-13c (20, 100; 21, 123; 22, 119; 23, 091). (24) Quarkus madstonensis Pessagno, Blome, and Hull in Pessagno et al., sample BMW-31 (24, 029). (25–26) Quarkus japonicus (Yao), sample BMW-13b (25, 026); sample BMW-31 (26, 067). (27–28) Praewilliriedellum convexum (Yao), sample BMW-13b (27, 027); sample BMW-28 (28, 038). (29–30) Quarticella sp. B, sample BMW-28 (29, 056; 30, 010). (31−37) Hemicryptocapsa yaoi (Kozur), sample BMW-13c (31, 116; 32, 112; 34, 118); sample EW-158 (33, 028); sample BMW-33 (35, 011); sample BMW-13b (36, 073); sample BMW-32 (37, 061). (38) Gongylothorax sp. A, sample BMW-34 (38, 019). (39−42), Gongylothorax favosus Dumitrica, sample BMW-32 (39, 058); sample BMW-30 (40, 045); sample BMW-34 (41, 025); sample BMW-33 (42, 036). (43–44) Williriedellum yahazuense (Aita), sample BMW-34 (43, 032); sample BMW-33 (44, 009). (45−48) Williriedellum sp. cf. W. formosum (Chiari, Marcucci, and Prela), sample BMW-28 (45, 049; 46, 033; 47, 058); sample BMW-13b (48, 067). (49) Williriedellum sp. B, sample BMW-13b (49, 053). (50–51) Williriedellum crystallinum Dumitrica, sample BMW-32 (50, 032); sample BMW-30 (51, 007). (52) Quarticella ovalis Takemura, sample BMW-28 (52, 034). (53) Hiscocapsa kodrai (Chiari, Marcucci, and Prela), sample BMW-13b (53, 030). (54–55) Quarticella sp. C, sample BMW-13b (54, 041); sample BMW-28 (55, 050). (56), Quarticella cicciona (Chiari, Marcucci, and Prela), sample BMW-13b (56, 033). (57) Quarticella sp. D, sample BMW-28 (57, 041). (58) Quarticella sp. A, sample EW-158 (58, 065).
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