O’Dogherty et al.—Jurassic radiolarians from the Eastern Alps Genus Praewilliriedellum Kozur, 1984 Type species.—Praewilliriedellum cephalospinosum Kozur, 1984.


Other species.—Stichocapsa? pseudoconvexa Kemkin and Taketani, 2004; Stichocapsa convexa Yao, 1979; Stichocapsa robusta Matsuoka, 1984; and Theocorys renzae Schaaf, 1981.


Emended diagnosis.—Praewilliriedellum is emended in order to allocate clear tetracyrtid forms (e.g., Stichocapsa robusta Matsuoka). Many representatives of this genus have an abdo- men partially encased in the postabdominal segment, making it hard to differentiate externally. The degree of encasement is nevertheless much lower than in Williriedellum and other genera assigned to Williriedellidae.


Occurrence.—Upper Aalenian to upper Barremian.


Praewilliriedellum convexum (Yao, 1979) Figure 9.27−9.28


1979 2003 2006


Stichocapsa convexa Yao, p. 35, pl. 5, figs. 14–16; pl. 6, figs. 1–7.


Stichocapsa convexa; Suzuki and Gawlick, p. 212, fig. 6.51.


Stichocapsa convexa;O’Dogherty et al., p. 441, pl. 6, fig. 35. [See for complete synonymy]


2007b Stichocapsa convexa; Gawlick et al., figs. 7.15, 8.32, 17.23, 18.10.


2008 2009


Praewilliriedellum convexum; Beccaro et al., pl. 2, fig. 26.


Stichocapsa convexa; Suzuki and Gawlick, p. 186, figs. 5.54A–B.


2013 Praewilliriedellum convexum; Chiari et al., fig. 13n.


Praewilliriedellum robustum (Matsuoka, 1984) Figure 10.45?, 10.46−10.47


1984


2003 2006


Stichocapsa robusta Matsuoka, p. 146, pl. 1, figs. 6–13; pl. 2, figs. 7–12.


Stichocapsa robusta; Suzuki and Gawlick, p. 213, fig. 5.44.


Stichocapsa robusta;O’Dogherty et al., p. 441, pl. 6, figs. 31–34. [See for complete synonymy]


2007b Stichocapsa robusta; Gawlick et al., fig. 8.34. 2008


Praewilliriedellum robustum;Beccaro et al., pl. 2, fig. 27. 2013 Praewilliriedellum robustum; Chiari et al., fig. 13p.


Genus Quarkus Pessagno, Blome, and Hull in Pessagno et al., 1993


Type species.—Quarkus madstonensis Pessagno, Blome, and Hull in Pessagno et al., 1993.


Other species.—Stichocapsa japonica Yao, 1979.


Occurrence.—Lower Bajocian to upper Callovian. A very similar specimen, but questionable to this genus, was found in upper Berriasian material of the Mariana Trench (Stichocapsa sp. 8 Matsuoka, 1998, pl. 2, fig. 24). We do not assign this species to Quarkus because continuous record from the Callovian to the Berriasian has not been demostrated yet.


Quarkus japonicus (Yao, 1979) Figure 9.25−9.26


1979 Stichocapsa japonica Yao, p. 36, pl. 6, figs. 8−12; pl. 7, figs. 1−15.


2006 Stichocapsa japonica;O’Dogherty et al., p. 441, pl. 6, fig. 6. [See for complete synonymy]


2009 Stichocapsa japonica; Suzuki and Gawlick, p. 186, figs. 5.53.


2012 Praewilliriedellum japonicum;Goričan et al., pl. 1, fig. 20. 2013 Praewilliriedellum japonicum; Chiari et al., fig. 13o.


Remarks.—This species has always been assigned to the genus Stichocapsa (considered as a nomen dubium by O’Dogherty et al., 2009, p. 330), however we note that this commonly illu- strated species is very close to Quarkus madstonensis, the only difference being the less-expanded fourth segment. Judging from the stratigraphic distribution, Quarkus japonicus may be the ancestor of Quarkus madstonensis. An intermediate form (Stichocapsa sp. aff. S. japonica Yao) was illustrated by Chiari et al., 2002 (pl. 3, figs. 18−22) from the upper Bajocian−lower Bathonian of Albania.


Quarkus madstonensis Pessagno, Blome, and Hull in Pessagno et al., 1993 Figure 9.24


1993 Quarkus madstonensis Pessagno, Blome, and Hull in Pessagno et al., p. 159, pl. 8, figs. 9, 13–15, 24.


2006 Williriedellum madstonense;O’Dogherty et al., p. 446, pl. 6, figs. 9–10.


2008 Williriedellum madstonense; Danelian et al., pl. 1, fig. 11.


Figure 10. Scale bars (a, b) measure 50µm; each radiolarian scanning micrograph is designated by sample and specimen number, followed by the corresponding scale. (1–2) Williriedellum? sp. C, sample BMW-13b (1, 066/a); sample BMW-28 (2, 007/a). (3) Arcanicapsa exquisita (Hull), sample BMW-32 (3, 013/a). (4) Arcanicapsa sp.A,sampleBMW-13b (4, 029/a). (5−16) Arcanicapsa funatoensis (Aita), sample BMW-26 (5, 035/a; 6, 031/a; 10, 037/a; 13, 035/a; 15, 032/a); sample EW-158 (7, 021/a; 8, 029/a; 9, 019/a); sample BMW-34 (11, 011/a); sample BMW-13b (12, 103/a); sample BMW-13c (14, 106/a; 16, 132/a). (17−19) Zhamoidellum ventricosum Dumitrica, sample BMW-13c (17, 121/a; 18, 114/a; 19, 136/a). (20−22) Arcanicapsa undulata (Heitzer), sample BMW-32 (20, 024/a); sample BMW-33 (21, 032/a); sample BMW-34 (22, 016/a). (23–24) Williriedellum sp.A,sample BMW-26(23, 059/a); sample BMW-13b (24, 074/b). (25−26) Kilinora sp. aff. K. oblongula (Kocher in Baumgartner et al.), sample EW-158 (25, 027/b); sample BMW-32 (26, 047/b). (27−30) Zhamoidellum sp. B, sample BMW-13c (27, 093/b); sample BMW-35 (28, 033/b); sample BMW-32 (29, 084/b); sample BMW-30 (30, 013/b). (31−40) Zhamoidellum ovum Dumitrica, sample BMW-33 (31, 021/a; 32, 012/a; 33, 016/a; 35,017/a; 39, 034/a); sample BMW-32 (34, 046/a); sample BMW-13c (36, 130/a; 37, 101/a); sample BMW-35 (38, 034/a); sample BMW-34 (40, 018/a). (41−44) Zhamoidellum sp.C,sample BMW-26(41, 005/a; 44, 063/a); sample BMW-35 (42, 035/a); sample EW-158 (43, 004/a). (45−47) Praewilliriedellum robustum (Matsuoka), sample EW-158 (?45, 025/b); sample BMW-31 (46, 071/a); sample BMW-34 (47, 017/a).


43


Page 1  |  Page 2  |  Page 3  |  Page 4  |  Page 5  |  Page 6  |  Page 7  |  Page 8  |  Page 9  |  Page 10  |  Page 11  |  Page 12  |  Page 13  |  Page 14  |  Page 15  |  Page 16  |  Page 17  |  Page 18  |  Page 19  |  Page 20  |  Page 21  |  Page 22  |  Page 23  |  Page 24  |  Page 25  |  Page 26  |  Page 27  |  Page 28  |  Page 29  |  Page 30  |  Page 31  |  Page 32  |  Page 33  |  Page 34  |  Page 35  |  Page 36  |  Page 37  |  Page 38  |  Page 39  |  Page 40  |  Page 41  |  Page 42  |  Page 43  |  Page 44  |  Page 45  |  Page 46  |  Page 47  |  Page 48  |  Page 49  |  Page 50  |  Page 51  |  Page 52  |  Page 53  |  Page 54  |  Page 55  |  Page 56  |  Page 57  |  Page 58  |  Page 59  |  Page 60  |  Page 61  |  Page 62  |  Page 63  |  Page 64  |  Page 65  |  Page 66  |  Page 67  |  Page 68  |  Page 69  |  Page 70  |  Page 71  |  Page 72  |  Page 73  |  Page 74  |  Page 75  |  Page 76  |  Page 77  |  Page 78  |  Page 79  |  Page 80  |  Page 81  |  Page 82  |  Page 83  |  Page 84  |  Page 85  |  Page 86  |  Page 87  |  Page 88  |  Page 89  |  Page 90  |  Page 91  |  Page 92  |  Page 93  |  Page 94  |  Page 95  |  Page 96  |  Page 97  |  Page 98  |  Page 99  |  Page 100  |  Page 101  |  Page 102  |  Page 103  |  Page 104  |  Page 105  |  Page 106  |  Page 107  |  Page 108  |  Page 109  |  Page 110  |  Page 111  |  Page 112  |  Page 113  |  Page 114  |  Page 115  |  Page 116  |  Page 117  |  Page 118  |  Page 119  |  Page 120  |  Page 121  |  Page 122  |  Page 123  |  Page 124  |  Page 125  |  Page 126  |  Page 127  |  Page 128  |  Page 129  |  Page 130  |  Page 131  |  Page 132  |  Page 133  |  Page 134  |  Page 135  |  Page 136  |  Page 137  |  Page 138  |  Page 139  |  Page 140  |  Page 141  |  Page 142  |  Page 143  |  Page 144  |  Page 145  |  Page 146  |  Page 147  |  Page 148  |  Page 149  |  Page 150  |  Page 151  |  Page 152  |  Page 153  |  Page 154  |  Page 155  |  Page 156  |  Page 157  |  Page 158  |  Page 159  |  Page 160  |  Page 161  |  Page 162  |  Page 163  |  Page 164  |  Page 165  |  Page 166  |  Page 167  |  Page 168  |  Page 169  |  Page 170  |  Page 171  |  Page 172  |  Page 173  |  Page 174  |  Page 175  |  Page 176  |  Page 177  |  Page 178  |  Page 179  |  Page 180  |  Page 181  |  Page 182  |  Page 183  |  Page 184  |  Page 185  |  Page 186  |  Page 187  |  Page 188  |  Page 189  |  Page 190  |  Page 191  |  Page 192  |  Page 193  |  Page 194  |  Page 195  |  Page 196  |  Page 197  |  Page 198  |  Page 199  |  Page 200  |  Page 201  |  Page 202  |  Page 203  |  Page 204  |  Page 205  |  Page 206  |  Page 207  |  Page 208