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Futakami and Haggart—early Albian (Early Cretaceous) douvilleiceratid ammonites


Table 3. Dimensions of specimens of Douvilleiceras offarcinatum (White, 1887) from Haida Gwaii, British Columbia


Specimen D H B B/H U U/D NR NT


GSC No. 10709 57.1 22.6 30.1 1.33 19.8 0.35 21 5 All measurements in millimeters.


Description.—The specimen GSC No. 10709 is approximately 57mmin diameter. The specimen has rectiradiate to very gently rursiradiate ribs and lacks intercalated ribs on the outer whorl. Ribs number 21 per whorl. On the outer whorl there are five rows of tubercles on each rib, consisting of three clavate tuber- cles in the ventrolateral region, a bullate or spinose lateral tubercle, and an umbilical tubercle, as shown in Figure 10.5. The medial ventral sulcus is wide.


Dimensions.—See Table 3.


Material.—A single specimen, GSC No. 10709, obtained from a sandy calcareous concretion at GSC Loc. C-304101, Haida Formation.


Occurrence.—This species is known from the upper lower Albian, the mammillatum Zone of the lower part of the Haida Formation in Canada, the Cuchillo Formation in Texas, and Sergipe Province in Brazil.


Remarks.—The morphological characters of this specimen are closely similar to those of the holotype of D. aurarium Anderson (1938, p. 175, pl. 53, fig. 1) from Shasta County, northern California. It is difficult to distinguish D. aurarium from D. offarcinatum (White, 1887) from Brazil, particularly in the mode and number of tubercles on both examples. Our specimen is somewhat similar to D. spiniferum; a distinct difference between them, however, is that our example of D. offarcinatum shows no intermediate tubercle between the lateral and umbilical tubercles, even in the middle growth stage at 57mm in diameter. Because the intermediate tubercles in D. spiniferum are already present on the flanks at this growth stage, the characteristic tuberculation of this specimen thus appears to fall outside the range of variation of D. spiniferum.


Douvilleiceras scabrosum Casey, 1962 Figures 10.1, 10.2, 11


? 1841 Ammonites mammillaris;d’Orbigny, p. 249, pl. 72, fig. 5.


1962 Douvilleiceras scabrosum; Casey, p. 278, pl. 40, figs. 1–3, pl. 42, figs. 2a-b, text-figs. 102f, 103i.


Types.—The holotype, BM C69861, illustrated by Casey (1962, pl. 40, figs. 1a-b, text-fig. 102f), from Folkstone Beds, mammillatum Zone Bed, Chrismill Bridge (M20 Motorway), Eyhorne Street, Kent (Coll. H. G. Owen); paratypes GSM 107909 (Casey, 1962, pl. 40, figs. 2a-b), GSM 70424 (Casey, 1962, pl. 40, figs. 3a-b, text-fig. 103i), GSM 107913 (Casey, 1962, pl. 42, figs. 2a-b), Folkestone Beds, main mammillatum


1972 Douvilleiceras sp. a; McLearn, p. 67, pl. 12, figs. 1A, B.


53


Table 4. Dimensions of specimens of Douvilleiceras scabrosum Casey, 1962 from Haida Gwaii, British Columbia


Specimen GSC No. 21241* 139.0 54.8 -


D H B B/H U U/D NR NT - 46.5 0.33 38 7 +1**


Notes All measurements in millimeters. *Measurements taken from plaster cast. **Indicating an additional row on each side of the venter.


bed, Copt Point, Folkestone, Kent (Coll. R. Casey), GSM Zn940 from the Folkestone Beds, the puzosianus Subzone, Sandling Junction, and BM C54304 from Machéroménil, Ardennes.


Dimensions.—See Table 4.


Material.—A single specimen, GSC No. 21241, described and illustrated by McLearn (1972, p. 67, pl. 12, figs. 1A-B; illustrated herein in Fig. 10.1, 2) as Douvilleiceras sp. a, is from GSC Loc. 7594.


Occurrence.—This species is known from the mammillatum Zone of the lower part of the Haida Formation in Canada and the Lower Greensand of southern England and France.


Remarks.—This specimen was identified by McLearn (1972, p. 67, pl. 12, figs. 1A-B) as Douvilleiceras sp. a. McLearn described the features of this specimen, giving comparisons with two other species of Douvilleiceras, D. spiniferum and D. scabrosum. McLearn concluded that the specimen either represents an extreme variant of D. spiniferum or a new species. We re-examined the specimen GSC No. 21241. The ribs are unequal in strength and rarely intercalated on the outer whorl. The tuberculation is represented by four distinct clavate tubercles in the ventrolateral region, a strong lateral spinose tubercle, a faint intermediate node-like tubercle, and a bullate umbilical one (Fig. 11.1). Based on the composition of these tubercles, this specimen falls distinctly outside of the range of variation of D. spiniferum. The morphological features of this specimen, particularly


the mode of tuberculation, the number of rows of tubercles on each side, and the mode of ribbing, are closely similar to those of the paratype of D. scabrosum Casey, GSM 107909 (Casey, 1962, p. 278, pl. 40, figs. 2a-b), from the mammillatum Zone of England. The Canadian specimen has somewhat weaker ribs than the paratype of D. scabrosum, and the initial appearance of the intermediate tubercles in this specimen is somewhat later than in the holotype and paratypes of D. scabrosum. It does appear, however, to fall within the relatively wide variation of D. scabrosum. According to Casey (1962, p. 278), D. scabrosum differs from D. mammillatum in having irregular and coarse ribbing and a wide ventral sulcus. In addition, the number of rows of tubercles on each side is seven, although the holotype specimen exhibits an additional row


of tubercles on each side of the venter (Casey, 1962, p. 285, text- fig. 102f).


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