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G President


OFFICERS AND EDITORS OF THE PALEONTOLOGICAL SOCIETY JOSEPH HANNIBAL,


MARY SILCOX,


STEVE HOLLAND, Univ. of Georgia stratum@uga.edu


President-Elect


ARNOLD I. MILLER, Univ. of Cincinnati


Arnold.miller@uc.edu Past President


SANDRA J. CARLSON,


Univ. of California–Davis sjcarlson@ucdavis.edu


Secretary


MARGARET M. YACOBUCCI, Bowling Green State Univ. mmyacob@bgsu.edu


Treasurer


PETER J. HARRIES, Univ. of South Florida


harries@chuma.cas.usf.edu JP Editors


BRENDA R. HUNDA Cincinnati Museum Center BHunda@cincymuseum.org


JISUO JIN


Univ. of Western Ontario jjin@uwo.ca


JP Managing Editor SARA A. MARCUS


journalofpaleontology@cambridge.org JP Associate Editors


JUAN LUIS BENEDETTO, Univ. Nacional de Cordoba jbenedetto@efn.uncor.edu


NICHOLAS BUTTERFIELD, Cambridge Univ.


njb1005@esc.cam.ac.uk


STYLIANOS CHATZIMANOLIS, Univ. Tennessee–Chattanooga stylianos-chatzimanolis@utc.edu


TOM CRONIN, USGS tcronin@usgs.gov


WILLIAM A. DIMICHELE, Smithsonian Institution dimichel@si.edu


ROBERT J. ELIAS, Univ. of Manitoba


eliasrj@ms.umanitoba.ca NADIA FRÖBISCH,


Museum für Naturkunde


nadiafroebisch@mfn-berlin.de FOREST J. GAHN,


Brigham Young Univ., Idaho gahnf@byui.edu


Cleveland Museum of Natural History jhannibal@cmnh.org


PETER J. HARRIES, Univ. of South Florida


harries@chuma.cas.usf.edu


JOACHIM HAUG, Univ. of Greifswald


joachim.haug@palaeo-evo-devo.info


MICHAEL HAUTMANN, Univ. of Zurich


michael.hautmann@pim.uzh.ch


MARTIN J. HEAD, Brock Univ.


mjhead@brocku.ca


SAMANTHA HOPKINS, Univ. of Oregon


shopkins@uoregon.edu


NIGEL HUGHES, UC Riverside


nigel.hughes@ucr.edu


DANIEL KSEPKA, Bruce Museum


dksepka@brucemuseum.org MAX LANGER,


Universidade de São Paulo mclanger@ff clrp.usp.br


STEPHEN A. LESLIE, James Madison Univ. lesliesa@jmu.edu


BRUCE LIEBERMAN, Univ. of Kansas blieber@ku.edu


STEPHEN LoDUCA,


Eastern Michigan Univ. sloduca@emich.edu


RICK LUPIA, OK Museum rlupia@ou.edu


GABRIELA MÁNGANO, Univ. Saskatchewan


gabriela.mangano@usask.ca


MALGORZATA MOCZYDLOWSKA- VIDAL, Uppsala Univ. malgo.vidal@pal.uu.se


ALISON MURRAY, Univ. of Alberta


ammurray@ualberta.ca


PAULA NOBLE, Univ. of Nevada noblepj@unr.edu


ERIC SCOTT,


San Bernardino Co. Museum escott@sbcm.sbcounty.gov


Univ. of Toronto Scarborough msilcox@utsc.utoronto.ca


HANS-DIETER SUES,


Smithsonian Institution suesh@si.edu


COLIN SUMRALL, Univ. of Tennessee csumrall@utk.edu


LEIF TAPANILA, Idaho State Univ. tapaleif@isu.edu


DAVID VARRICCHIO, Montana State Univ. djv@montana.edu


PETER WAGNER,


Smithsonian Institution wagnerpj@si.edu


STEVE WESTROP, OK Museum


swestrop@ou.edu DAVID M. WORK,


Maine State Museum david.work@maine.gov


PATRICK WYSE JACKSON, Trinity College Dublin wysjcknp@tcd.ie


MARGARET M. YACOBUCCI, Bowling Green State Univ. mmyacob@bgsu.edu


SAMUEL ZAMORA,


Geological Survey of Spain samuel@unizar.es


RENBIN ZHAN,


Nanjing Institute of Geology and Palaeontology,


Chinese Academy of Sciences rbzhan@nigpas.ac.cn


Paleobiology Co-Editors


MICHAEL KOWALEWSKI Univ. of Florida


CATHERINE BADGLEY Univ. of Michigan


SHUHAI XIAO Virginia Tech


CARLOS A. JARAMILLO


Smithsonian Tropical Research Institute


Editor of Special Publications SARA A. MARCUS,


Oregon Inst. Of Technology sara.marcus@oit.edu


Communications Offi cer LEIF TAPANILA, Idaho State Univ. tapleif@isu.edu


G


Program Coordinator


MATTHEW CLAPHAM Univ. of California, Santa Cruz


mclapham@ucsc.edu


Education/Outreach Coordinator MARGRET FRAISER,


Univ. of Wisconsin, Milwaukee mfraiser@uwm.edu


Councilors LAUREL COLLINS,


Florida International Univ. collinsl@fi u.edu


KATE BULINSKI, Bellarmine University kbulinski@bellarmine.edu


Student Representative LUCY CHANG,


Univ. California, Berkeley luchang@berkeley.edu


SHARON McMULLEN,


Univ. of Wisconisin–Madison skmcmullen@wisc.edu


Section Chairs Cordilleran:


NICOLE BONUSCO, CA State Univ. Fullerton nbonuso@fullerton.edu


North Central:


REBECCA FREEMAN, Univ. of Kentucky


rebecca.freeman@uky.edu Northeastern:


ALEXANDER J. BARTHOLOMEW, SUNY–New Paltz


barthola@newpaltz.edu


Rocky Mountain: LEIF TAPANILA, Idaho State Univ. tapaleif@isu.edu


Southeastern:


CHRISTY VISAGGI, Georgia State Univ. cvisaggi@gsu.edu


Editor – Priscum MATT POWELL, Juniata College


powell@juniata.edu


Editor – Social Media PHOEBE COHEN, Williams College


Phoebe.A.Cohen@williams.edu


Cover illustration: T e early Oligocene cetacean, Agorophius pygmaeus, type specimen, reproduced from the plate of Phocodon holmesii Agassiz, from Godfrey et al., Fig 1.3, p. 155.


ISSN 0022-3360 Copyright © 2016 T e Paleontological Society. All rights reserved.


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