180
Journal of Paleontology 90(1):170–181
and grazing herbivores and it is likely that Bothriogenys focused on foliage. In particular, given the apparent hydrophyllic nature of some anthracotheres (Pickford, 2008), Bothriogenys might have relied on aquatic plants as dietary staples. Nabotherium,in contrast, with its relatively shorter rostrum, enlarged projecting canines, short to absent anterior diastemata, low-crowned and simple premolars, and low, broad, bunodont molars, was likely to have been a more eclectic frugivore/herbivore.
Acknowledgments
We thank O. Teamaa, chairman of Egyptian Mineral Resources Authority, S. Mansour, director of the Central Administration at the Egyptian Geological Survey, R. Hraoun, general director of Egyptian Geological Museum, and M. Abdel Ghani, curator of invertebrate Paleontology Collections in the Egyptian Geological Museum, for facilitating this work. We would like to thank S. El-Sayed and I. El Dawoudi from Mansoura University Vertebrate Paleontology Center (MUVP) for assistancewith preparation and other aspects of the research. Two anonymous reviewers helped to improve a previous version of this paper.
References
Andrews, C.W., 1906, A descriptive catalogue of the Tertiary Vertebrata of the Fayum, Egypt. Based on the collection of the Egyptian government in the Geological Museum, Cairo, and on the collection in the British Museum (Natural History), London: London,BritishMuseumofNatural History, 324 p.
Colbert, E.H., 1938, Fossil Mammals from Burma in the American Museum of Natural History: Bulletin of the American Museum of Natural History, v. 74, p. 255–436.
Cuvier, G., 1822, Recherches sur les Ossemens Fossils, où l’on Rètabit les Caractères de Plusieurs Animax don’t les Revolutions du Globe ont Détruit les Espèces: Paris, Dufour et E. d’Ocagne, 412 p.
Domning, D.P., and Gingerich, P.D., 1994, Protosiren smithae, new species (Mammalia, Sirenia), from the late middle Eocene of Wadi Hitan, Egypt: Contributions from the Museum of Paleontology, University of Michigan, v. 29, p. 69–87.
Domning, D.P., Gingerich, P.D, Simons, E.L., and Ankel-Simons, F.A., 1994, A new early Oligocene dugongid (Mammalia, Sirenia) from Fayum Province, Egypt: Contributions from the Museum of Paleontology, University of Michigan, v. 29, p. 89–108.
Ducrocq, S., 1997, The anthracotheriid genus Bothriogenys (Mammalia, Artio- dactyla) in Africa and Asia during the Paleogene: Phylogenetical and paleobiogeographical relationships: Stuttgarter Beitrage zur Naturkunde, B (Geologie und Palaontologie), v. 250, p. 1–44.
Erfurt, J., and Sudre, J., 1995, Un Haplobunodontidae nouveau, Hallebune krumbiegeli nov. gen. nov. sp. (Artiodactyla, Mammalia), dans l’Eocene moyen du Geiseltal pres halle (Sachsen-Anhalt, Allemagne): Palaeo- vertebrata, v. 24, p. 84–99.
Gingerich, P.D., Smith, B.H., and Simons, E.L., 1990, Hind limbs of Eocene Basilosaurus isis: Evidence of feet in whales: Science, v. 249, p. 154–157.
Holroyd, P., Lihoreau, P., Gunnell, G.F., and Miller, E.R., 2010, Anthra- cotheriidae, in Werdelin, L., and Sanders, W.J., eds., Cenozoic Mammals of Africa: Berkeley, University of California Press, p. 843–851.
Hooker, J.J., and Thomas, K.M., 2001, A new species of Amphirhagatherium (Choeropotamidae, Artiodactyla, Mammalia) from the late Eocene Headon Hill Formation of southern England and phylogeny of endemic European ‘anthracotherioids’: Palaeontology, v. 44, p. 827–853.
Khin, Zaw, Meffre, S., Takai, M., Suzuki, H., Burrett, C., Thaung, Htike, Zin, Maung, Maung, Thein, Tsubamoto, T., Egi, N., and Maung Maung, 2014, The oldest anthropoid primates in SE Asia: Evidence from LA-ICP-MSU- Pb zircon age in the late middle Eocene Pondaung Formation, Myanmar: Gondwana Research, v. 26, p. 122–131.
Kowalevsky, W., 1874, Monographie der Gattung Anthracotherium: Palaeon- tographica, v. 22, p. 131–290.
Leidy, J., 1869, The extinct mammalian fauna of Dakota and Nebraska, including an account of some allied forms from other localities, together with a synopsis of the mammalian remain of North America: Journal of the Academy of Natural Sciences, Philadelphia, ser. 2, v. 7, p. 1–472.
Lihoreau, F., and Ducrocq, S., 2007, FamilyAnthracotheriidae, in Prothero, D.R., and Foss, S.E., eds., The Evolution of Artiodactyls, Baltimore, Maryland, The Johns Hopkins University Press, p. 89–105.
Lihoreau, F., Boisserie, J.R., Manthi, F.K., and Ducrocq, S., 2015, Hippos stem from the longest sequence of terrestrial cetartiodactyl evolution in Africa: Nature Communications, v. 6, p. 6264, doi:10.1038/ncomms7264.
Linnaeus, C., 1758, Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentis, Synonymis, Locis, 10th ed., Stockholm, Sweden, Laurentii, Slavi, 824 p.
Macdonald, J.R., 1956, The North American anthracotheres: Journal of Paleontology, v. 30, p. 615–645.
McKenna, M.C., and Bell, S.J., 1997, Classification of Mammals Above the Species Level: New York, Columbia University Press, 631 p.
Murray, A.M., Cook, T.D., Attia, Y.S., Chatrath, P., and Simons, E.L., 2010, A freshwater ichthyofauna from the late Eocene Birket Qarun Formation, Fayum, Egypt: Journal of Vertebrate Paleontology, v. 30, p. 665–680.
Owen, R., 1848, Description of teeth and portions of two of two extinct anthracotheroid quadrapeds (Hyopotamus vectianus and H. bovinus) dis- covered by the Marchioness of Hastings in the Eocene deposits on the N.W. coast of the Isle of Wight, with an attempt to develop Cuvier’s idea of the classification of pachyderms by the number of their toes: Quarterly Journal of the Geological Society of London, v. 4, p. 104–141.
Pickford, M., 2008, Libycosaurus petrocchii Bonarelli, 1947, and Libycosaurus anisae (Black, 1972) (Anthracotheriidae, Mammalia): Nomenclatural and geochronological implications: Annales de Paléontologie, v. 94, p. 39–55.
Pictet, F.J., 1855–1857, Mémoire sur les Animaux Vertébrés Trouvés dans le Terrain Sidérolitique du Canton de Vaud et Appartenant a la Faune Éocene, Seconde Partie, Description des Ossements Fossiles: Geneva, Switzerland, J. Kessman, Libraire, p. 27–120.
Pictet, F.J., 1857, Seconde Partie. Description des ossements fossiles trouvés au Mauremont, in Pictet, F.-J., Gaudin, C., and De La Harpe, P. 1855–1857, Mémoire sur les Animaux Vertébrés Trouvés dans le Terrain Sidérolithique du Canton de Vaud et Appartenant à la Faune Éocène, Volume 2, Matériaux de Paléontologie Suisse (note 1), Geneva, Switzerland, J. Kessmann, p. 27–120.
Pilgrim, G.E., 1928, The Artiodactyla of the Eocene of Burma: Palaeontologia Indica, v. 13, p. 1–39.
Pilgrim, G.E., 1941, The dispersal of the Artiodactyla: Biological Reviews, v. 16, p. 134–163.
Pilgrim, G.E., and Cotter, G.D., 1916, Some newly discovered Eocene mammals from Burma: Records of the Geological Survey of India, v. 47, p. 42–77.
Rasmussen, D.T., and Simons, E.L., 1988, New Oligocene hyracoids from Egypt: Journal of Vertebrate Paleontology, v. 8, p. 67–83.
Sallam, H.M., Seiffert, E.R., Steiper, M.E., and Simons, E.L., 2009, Fossil and molecular evidence constrain scenarios for the early evolutionary and bio- geographic history of hystricognathous rodents: Proceedings of the National Academy of Sciences, v. 106, p. 16722–16727.
Sallam, H.M., Seiffert, E.R., and Simons, E.L., 2011, Craniodental morphology and systematics of a new family of hystricognathous rodents (Gaudeamur- idae) from the late Eocene and early Oligocene of Egypt: PLoS ONE, v. 6, e16525.
Seiffert, E.R., 2006, Revised age estimates for the later Paleogene mammal faunas of Egypt and Oman: Proceedings of the National Academy of Sciences, v. 103, p. 5000–5005.
Seiffert, E.R., 2007, A new estimate of afrotherian phylogeny based on simul- taneous analysis of genomic, morphological, and fossil evidence: BMC Evolutionary Biology, v. 7, p. 224, doi:10.1186/1471-2148-7-224.
Seiffert, E.R., 2010, The oldest and youngest records of afrosoricid placentals from the Fayum Depression of northern Egypt: Acta Palaeontologica Polonica, v. 55, p. 599–610.
Seiffert, E.R., 2012, Early primate evolution in Afro-Arabia: Evolutionary Anthropology, v. 21, p. 239–253.
Seiffert, E.R., Simons, E.L., and Attia, Y., 2003, Fossil evidence for an ancient divergence of lorises and galagos: Nature, v. 422, p. 421–424.
Seiffert, E.R., Perry, J.M.G., Simons, E.L., and Boyer, D.M., 2009, Convergent evolution of anthropoid-like adaptations in Eocene adapiform primates: Nature, v. 461, p. 1118–1121.
Simons, E.L., and Rasmussen, D.T., 1990, Vertebrate paleontology of Fayum: History of research, faunal review and future prospects, in Said, R., ed., The Geology of Egypt: Rotterdam, The Netherlands, A. A. Balkema, p. 627–638.
Simons, E.L., Seiffert, E.R., Ryan, T.M., and Attia, Y., 2007, A remarkable female cranium of the early Oligocene anthropoid Aegyptopithecus zeuxis (Catarrhini, Propliopithecidae): Proceedings of the National Academy of Sciences, v. 104, p. 8731–8736.
Schmidt, M., 1913, Ueber Paarhufer der fluviomarinen Schichted des Fajum: Geologische und Paläontologische Abhandlungen, v. 15, p. 153–264.
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