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394 H. Somsen and A. Trouwborst


the CMS have also adopted treaty interpretations welcoming rather than deterring species driven upwards or polewards by climate change (e.g. CMS COP Resolution 12.21, 2017). In summary, current international wildlife law clearly


suggests that the coyote in Costa Rica, crab-eating fox in Panama, and golden jackal in the Netherlands are not to be considered as alien species, whether invasive or not. Thus, they are also not subject to any legal requirements to combat invasive alien species. It may be that measures to prevent damage inflicted by


canids on native species in newly colonized territories (e.g. coyotes eating Dice’s cottontails) are appropriate in certain situations, and this could conceivably include lethal control. Again, an evident role is reserved in this regard for inter- national or national wildlife law, under which colonizing ca- nids may be designated as, for example, protected, game or pest species. However, to regard the pioneering coyote, golden jackal and crab-eating fox as alien species to be dis- couraged or eradicated would be at odds with current inter- national wildlife law. Although our analysis is limited to clarifying the legal sta-


tus of colonizing versus introduced species, we draw attention to the existence of various related issues. For in- stance, further complications arise where range expansions into non-historical areas result from amixture of natural ex- pansion and anthropogenic introductions. An illustration is the red fox in lowland areas of North America, where the species’ establishment largely seems to have been a natural expansion, but was also influenced by introductions of foxes from Europe (Statham et al., 2012). Likewise, coyotes and golden jackals can hybridize with wolves and domestic dogs, raising vexing questions regarding the legal status of hybrid offspring (Trouwborst, 2014). These issues, along with de-extinction (Somsen, 2016) and assisted colonization (Trouwborst, 2015), are novel conundrums that wildlife law must come to grips with in the Anthropocene.


Acknowledgements AT was supported by the Netherlands Organization for Scientific Research, as part of the project Ius Carnivoris (No. 452-13-014). Helpful comments by two anonymous reviewers are gratefully acknowledged.


Author contributions Study design, legal analysis and writing the article: HS and AT.


Conflicts of interest None.


Ethical standards This research complied with the Oryx Code of Conduct.


References


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Oryx, 2020, 54(3), 392–394 © 2019 Fauna & Flora International doi:10.1017/S0030605318001229


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