Chen et al.—New Jurassic Sinoalidae from China


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(ZeissSteREO Discovery V8). Line drawings were prepared with CorelDraw 12.0 and Adobe Photoshop CS3.


Repository and institutional abbreviation.—All the material and figured specimens in this study are deposited in Shandong Tianyu Museum of Nature (STMN), Pingyi, Shandong Province, China.


Systematic paleontology


Order Hemiptera Linnaeus, 1758 Suborder Cicadomorpha Evans, 1946 Superfamily Cercopoidea Leach, 1815


Family Sinoalidae Wang and Szwedo in Wang et al., 2012


Figure 1. Locations of fossil specimens of the family Sinoalidae. 1, Daohugou, Ningcheng County, Inner Mongolia; 2, Zhouyingzi, Luangping County, Heibei.


updated International Chronostratigraphic Chart (Cohen et al., 2016).


Nel et al. (2012) proposed a new interpretation of wing


venation pattern for all Paraneoptera, assuming that CuA gets fused withM+R stem at the wing base and connected with CuP by a specialized crossvein cua-cup after its departure from M+R, which is remarkably different from the traditional interpretations. The venational terminologies used herein follow Nel et al. (2012). The fossil sinoalids were examined dry or under alcohol, with details observed and photographed under a stereomicroscope


Diagnosis (emended).—Forewing with apices of costal area and clavus almost at the same level; costal area and/or clavus more sclerotized and punctate and remaining parts membranous; Pc +CP long and thicken, almost parallel to costal margin; M two-branched. Hind wing withMunbranched; crossvein m-cua basad of crossvein r-m. Three ocelli rather than two. Hind tibia with two rows of lateral spines (four at most in number for each row).


Remarks.—Our new fossil materials suggest that the relative branching position of M and CuA is variable in Sinoalidae and so not appropriate for family-level diagnosis. In addition, the fossils reported herein also provide some information on morphological diversity and evolution of hind wings and hind tibiae of the family (see Discussion).


Key to genera of the family Sinoalidae on characters of forewing (updated from Wang et al. [2012]):


1. Crossvein m-cua connecting to stem M, bifurcation of M distad of apices of costal area and clavus, CuA branching into CuA1 and CuA2 at junction with crossvein m-cua, slightly basad of bifurcation ofM. . .... ..... .... .... ..... .... .... ..... .... .. Shufania n. gen. Crossvein m-cua absent or connecting toM3+4, bifurcation ofMbasad of apices of costal area and clavus, CuA branching into CuA1 and CuA2 far away from junction with crossvein m-cua, distad of bifurcation ofM ..... .... .... ..... .... .... ..... .... .... ..... .... . 2


2. ScP+RA ~1.3 times as long as stem ScP+R, CuA forking distad of apices of costal area, stigmal cell as wide as radial cell, crossvein m-cua absent.... .... .... ..... .... .... ..... .... .... ..... .... .... ..... .... .... ..... .... .... ..... Luanpingia Hong, 1983


ScP+RA at least 1.5 times as long as stem ScP+R, CuA forking basad of apices of costal area, stigmal cell about half as wide as radial cell, crossvein m-cua present . . . .... .... ..... .... .... ..... .... .... ..... .... .... ..... .... .... ..... .... .... ..... .... . 3


3. Apex truncate, basal cell ~0.12 times as long as forewing length, ScP+RA ~10 times as long as stem ScP+R, crossvein m-cua connecting to branch CuA1 .. .... ..... .... .... ..... .... .... ..... .... .... ..... .... .... Sinoala Wang and Szwedo in Wang et al., 2012


Apex rounded; basal cell long, ScP+RA at most 4.5 times as long as stem ScP+R, crossvein m-cua connecting to stem CuA ..... .... . 4


4. StemsScP,R,Mand CuAleavingbasal cellatcommonpoint,branchScP+RA~4.4times as long as ScP+ R.... Huabeicercopis Hong, 1983 Common stalk ScP+R + M existing, branch ScP+RA ~1.5 times as long as ScP +R .... . Jiania Wang and Szwedo in Wang et al., 2012


Genus Shufania new genus


Type species.—Shufania hani n. gen. n. sp., by present designation and monotypy.


Diagnosis.—As for the type species.


Etymology.—The generic name, Shufania, a feminine noun derived after Prof. Shufan Han, a well-known artist and the curator of the Museum of Linyi University.


Shufania hani new species Figure 2


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