194 J. Bonnald et al. References
ABDI,H.&WILLIAMS, L.J. (2010) Principal component analysis.Wiley Interdisciplinary Reviews: Computational Statistics, 2, 433–459.
BEDETTI, A., GREYLING, C., PAUL, B., BLONDEAU, J., CLARK, A., MALIN, H. et al. (2020) System for Elephant Ear-pattern Knowledge (SEEK) to identify individual African elephants. Pachyderm, 61, 63–77.
BONNALD, J., UTGE, J., KUHNER, M.K.,WASSER, S.K., ASALU, E., OKIMAT, J.P. & KRIEF,S.(2021) Who are the elephants living in the hybridization zone? How genetics may guide conservation to better protect endangered elephants. Global Ecology and Conservation, 32,e01917.
BORTOLAMIOL, S., COHEN, M., POTTS, K., PENNEC,F., RWABURINDORE, P., KASENENE, J. et al. (2014) Suitable habitats for endangered frugivorous mammals: small-scale comparison, regeneration forest and chimpanzee density in Kibale National Park, Uganda. PLOS ONE, 9,e102177.
CAPELOT,J. (2013) Des techniques de défenses entre traditions et innovations dans les conflits hommes-éléphants. Étude ethno-éthologique dans le nord-est du parc national de Kibale (Ouganda). Mémoire de Master 2 Spécialité Environnement, Développement, Territoires, Société, Muséum National d’Histoire Naturelle, Paris, France.
CHACÓN, J.E. &DUONG,T.(2018) Multivariate Kernel Smoothing and Its Applications. CRC Press, Boca Raton, USA.
CHAPMAN, C.A. & LAMBERT, J.E. (2000) Habitat alteration and the conservation of African primates: case study of Kibale National Park, Uganda. American Journal of Primatology, 50, 169–185.
COMSTOCK, K., GEORGIADIS, N., PECON-SLATTERY, J., ROCA, A., OSTRANDER, E., O’BRIEN,S. &WASSER, S.K. (2002) Patterns of molecular genetic variation among African elephant populations. Molecular Ecology, 11, 2489–2498.
DEBRUYNE,R. (2003) Différenciation morphologique et moléculaire des Elephantinae (Mammalia, Proboscidea). Statut systématique de l’éléphant d’Afrique de forêt, Loxodonta africana cyclotis (Matschie, 1900). Doctoral thesis. Muséum National d’Histoire Naturelle, Paris, France.
ELDER,W.H. (1970) Morphometry of elephant tusks. Zoologica Africana, 5, 143–159.
FALLON,F. (1944) L’Éléphant Africain. Publications de l’Institut Royal Colonial Belge, Section des sciences naturelles et médicales, Tome XIII, fasc. 2, Brussells, Belgium.
FOWLER, M.E. & MIKOTA, S.K. (2006) Biology, Medicine and Surgery of Elephants. Blackwell Publishing, Hoboken, USA.
GOBUSH, K.S., EDWARDS, C.T.T., BALFOUR, D.,WITTEMYER, G., MAISELS,F. & TAYLOR, R.D. (2021a) Loxodonta
africana.In The IUCN Red List of Threatened Species 2021.
dx.doi.org/10.2305/IUCN. UK.2021-2.RLTS.T181008073A204401095.en.
GOBUSH, K.S., EDWARDS, C.T.T., MAISELS, F.,WITTEMYER, G., BALFOUR,D.&TAYLOR, R.D. (2021b) Loxodonta
cyclotis.In The IUCN Red List of Threatened Species 2021.
dx.doi.org/10.2305/IUCN. UK.2021-1.RLTS.T181007989A204404464.en.
GROVES, C.P. (2000) What are the elephants ofWest Africa? Elephant, 2, 7–8.
GROVES, C.P. & GRUBB,P.(2000)Do Loxodonta cyclotis and L. africana interbreed? Elephant, 2, 4–7.
GRUBB, P., GROVES, C.P, DUDLEY, J.P. & SHOSHANI,J.(2000) Living African elephants belong to two species: Loxodonta africana (Blumenbach, 1797) and Loxodonta cyclotis (Matschie, 1900). Elephant, 2, 1–4.
HANKS,J.(1972) Growth of the African elephant (Loxodonta africana). African Journal of Ecology, 10, 251–272.
HART, J., GOBUSH, K., MAISELS, F.,WASSER, S., OKITA-OUMA,B.& SLOTOW,R.(2021) African forest and savannah elephants treated as separate species. Oryx, 55, 170–171.
HARTIGAN, J.A. &WONG, M.A. (1979) Algorithm AS 136:a k-means clustering algorithm. Journal of the Royal Statistical Society, 28, 100–108.
HOULE, D., GOVINDARAJU, D.R. & OMHOLT,S.(2010) Phenomics: the next challenge. Nature Reviews Genetics, 11, 855–866.
ISHIDA, Y., DEMEKE,Y., VAN COEVERDEN DE GROOT, P.J., GEORGIADIS, N.J., LEGGETT, K.E., FOX, V.E. & ROCA, A.L. (2011) Distinguishing forest and savanna African elephants using short nuclear DNA sequences. Journal of Heredity, 102, 610–616.
KEIGWIN, M.,WABUKAWO, V.,WASSER, S.K. & CHAPMAN,C.(2016) Impacts on transboundary elephant movements between Queen Elizabeth Conservation Area, Uganda and Parc National des Virunga, Democratic Republic of Congo. Pachyderm, 57, 118–121.
KIM, H.J. &WASSER, S.K. (2019) Loxodonta africana Subspecies Distribution Across African Elephant Database Input Zones. University of Washington, Seattle, USA.
iucn.org/sites/dev/files/ content/documents/2019-03-15-final-taxanomy_report- africanelephant-sg.pdf [accessed 13 April 2022].
LAWS, R.M. (1966) Age criteria for the African elephant, Loxodonta a. africana. African Journal of Ecology, 4, 1–37.
LAYSER, T.R. & BUSS, I.O. (1985) Observations on morphological characteristics of elephant tusks. Mammalia, 49, 407–414.
LÊ, S., JOSSE,J. & HUSSON,F.(2008) FactoMineR:an R package for multivariate analysis. Journal of Statistical Software, 25, 1–18.
MAISELS, F., STRINDBERG, S., BLAKE, S.,WITTEMYER, G., HART, J., WILLIAMSON, E.A. et al. (2013) Devastating decline of forest elephants in Central Africa. PLOS ONE, 8,e59469.
MONDOL, S., MOLTKE, I., HART, J., KEIGWIN, M., BROWN, L., STEPHENS,M.&WASSER, S.K. (2015) New evidence for hybrid zones of forest and savanna elephants in central and west Africa. Molecular Ecology, 24, 6134–6147.
MORRISON-SCOTT, T.C.S. (1947) A revision of our knowledge of African elephant’s teeth, with notes on forest and ‘pygmy’ elephants. Proceedings of the Zoological Society of London, 117, 505–527.
OMEJA, P.A., LAWES, M.J., CORRIVEAU, A.,VALENTA, K., SARKAR, D., PAIM, F.P. & CHAPMAN, C.A. (2016) Recovery of tree and mammal communities during large-scale forest regeneration in Kibale National Park. Biotropica, 48, 770–779.
PEDREGOSA, F., VAROQUAUX, G., GRAMFORT, A., MICHEL,V., THIRION, B., GRISEL, O. et al. (2011) Scikit-learn: machine learning in Python. Journal of Machine Learning Research, 12, 2825–2830.
PFEFFER,P.(1989) Vie et Mort d’un Géant: L’Éléphant d’Afrique. Flammarion, Paris, France.
RCORE TEAM (2019) R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, Vienna,Austria.
RIPLEY, B.D. (2007) Pattern Recognition and Neural Networks. Cambridge University Press, Cambridge, UK.
ROCA, A.L., GEORGIADIS,N.&O’BRIEN, S.J. (2005) Cytonuclear genomic dissociation in African elephant species. Nature Genetics, 37, 96–100.
ROCA, A.L., GEORGIADIS, N., PECON-SLATTERY,J. & O’BRIEN, S.J. (2001) Genetic evidence for two species of elephant in Africa. Science, 293, 1473–1477.
ROTH, H.H.&DOUGLAS-HAMILTON,I. (1991) Distribution and status of elephants in West Africa. Mammalia, 55, 489–527.
SHOSHANI,J.(1993) How many species or subspecies of forest Loxodonta exist? In Abstracts, Sixth International Theriological Congress, pp. 278–279. 4–10 July 1993, Sydney, Australia.
SIKES, S.K. (1966) The African elephant, Loxodonta africana:a field method for the estimation of age. Journal of Zoolology, 150, 279–295.
Oryx, 2023, 57(2), 188–195 © The Author(s), 2022. Published by Cambridge University Press on behalf of Fauna & Flora International doi:10.1017/S0030605321001605
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