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Fossil Weathering and Preparation Mimic Soft Tissues


operated by the National Research Agency (ANR). Te authors thank Stefan Lalonde and Pierre Sansjofre (Brest University) for assistance during SEM and XRF analyses. Te Editor-in-Chief, Bob Price, is thanked for his helpful and constructive remarks.


References [1] P Van Roy et al., Nature 465 (2010) 215–18. [2] P Van Roy et al., J Geol Soc 172 (2015) 541–49. [3] B Lefebvre et al., Palaeogeogr Palaeoclimatol Palaeoecol 460 (2016) 97–121.


[4] P Van Roy and DEG Briggs, Nature 473 (2011) 510–13. [5] JC Gutiérrez-Marco et al., Sci Rep 7 (2017) 39728. [6] J Vinther et al., Nature 451 (2008) 185–88. [7] M Marti Mus, Palaeogeogr Palaeoclimatol Palaeoecol 460 (2016) 122–29.


Figure 4: (A–B) Carbon particles (black) in tube feet-like structures of the putative water vascular system, (C–F) carbon particles found in other random structures showing the same carbon distribution as tube feet-like structures.


of putative soſt parts in eocrinoids and somasteroids from the Fezouata Shale [20,21] are either preparation artifacts or the result of recent weathering. In addition, it shows that under- standing the history of a specific fossil, from the excavation to preparation, is essential for proper paleontological interpreta- tion, especially when it comes to the description of soſt tissues in extinct taxa. Tese results should be taken into consider- ation when studying any new fossil, whether it is purchased from a private collector or comes from a museum collection.


Acknowledgements Tis paper is a contribution to the TelluS-INTERRVIE proj-


ects “Mécanismes de préservation exceptionnelle dans la Forma- tion des Fezouata” (2018), and “Géochimie d’un Lagerstätte de l’Ordovicien inférieur du Maroc” (2019), all funded by the INSU (Institut National des Sciences de l’Univers, France), CNRS. Tis paper is also a contribution to the International Geoscience Pro- gramme (IGCP) Project 653 – Te Onset of the Great Ordovician Biodiversification Event. Te authors are grateful to the LABEX Lyon Institute of Origins (ANR-10-LABX-0066) of the Université de Lyon for its financial support within the program “Investisse- ments d’Avenir” (ANR-11-IDEX-0007) of the French government


[8] J Vinther et al., Nature 542 (2017) 471–74. [9] ELO Martin et al., Palaeogeogr Palaeoclimatol Palaeoecol 460 (2016) 130–41.


[10] K Kouraïss et al., Palaeogeogr Palaeoclimatol Palaeoecol 508 (2018) 48–58.


[11] B Lefebvre et al., Progr. Abstr. 57th 44–45.


Pal’Ass Meeting (2013)


[12] B Lefebvre et al., Geobios 52 (2018) 27–36. [13] MD Sutton et al., Proc R Soc B 272 (2005) 1001–06. [14] DEG Briggs et al., Proc R Soc B 284 (2017) 20171189. [15] IA Rahman et al., Proc R Soc B 286 (2019) 20182792. [16] A Glass and DB Blake, Paläont Z 78 (2004) 73–95. [17] CE Brett et al., Paleont Soc Papers 3 (1997) 147–90. [18] P Gorzelak and MA Salamon, Palaeogeogr Palaeoclimatol Palaeoecol 386 (2013) 569–74.


[19] F Saleh et al., Earth Planet Sc Lett 529 (2020) 115873. [20] B Lefebvre et al., Progr. Abstr. 62nd [21] M Nohejlová, Abstr., 5th


Pal’Ass Meeting (2018) 40. Int. Palaeont. Congr. (2018) 851.


[22] ELO Martin, Communautés animales du début de l’Ordovicien (∼480 Ma): etudes qualitatives et quantitatives à partir de sites à preservation exceptionnelle des Fezouata, Maroc, Unpubl. PhD thesis, Lyon, 2016, pp. 1–313.


[23] F Saleh et al., Geology 47 (2018) 103–06. [24] R Vaucher et al., Sedimentology 64 (2017) 777–807. [25] SE Gabbott et al., Geology 32 (2004) 901–04. [26] RR Gaines et al., Geology 36 (2008) 755–58.


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