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5 the state of science in south africa
B I O L O G I C A L S C I E N C E S
suitable for forests, and it is probably fre- The costs and benefits of changes in woody
quent fires that restrain the growth of cover in the tropical grassy biomes vary, de-
woody vegetation. Mean annual rainfall, pending on ecological and socio-economic
and fire, are the most important correlates contexts, and spatial scale. Carbon seques-
of tree cover, with African megafauna, par- tration projects have become a source of in-
ticularly elephants (where they still exist) come in some regions where increased tree
contributing to open grassy structure. The cover is a desirable goal. Elsewhere, tree in-
advent of satellite imagery has led to an crease is undesirable, as in African savanna
increased appreciation of fire in impacting parks and livestock farming areas. Forest
on both vegetation structure and global cli- spread into grasslands could cause casca-
mate change. ding changes in biodiversity. Holistic studies
of the consequences of changes in savanna
The future of savannas and grasslands is landscapes are needed to evaluate fully the
uncertain. The direct effects of increasing social and economic impacts on society
atmospheric CO
2
are likely to be consider- and on nature; this is an area that requires
able, given that C3 woody vegetation is further directed research in South Africa.
likely to respond more than the C4 grasses.
A consequence of this is that woody spe- Grassland science
cies may grow sufficiently rapidly to escape Grassland science is an applied form of
the flame zone and thus become well es- plant science in South Africa referring to
tablished. The overall effect of increasing the ecology and management of natural
CO
2
in grassy ecosystems is to promote grassland and cultivated pasture. The term
woody plants within savannas, leading to can be somewhat misleading in that grass-
increased woody cover. Indeed, in South dominated vegetation occurs in many of
Africa, scrub forest has invaded savannas the biomes (other than the grassland bi-
in conservation, commercial ranching and ome) occurring in South Africa. Internation-
communally farmed areas. Acting against ally, ‘rangeland’ is a more commonly used
CO
2
-induced increases in woody cover are term to describe natural, grass-dominated
climate-induced increases in tree mortality vegetation.
from drought or high-severity burns.
In South Africa, grassland science originat-
Changes in woody cover have many con- ed out of a need to understand and man-
sequences, both positive and negative, age the vast natural veld used primarily for
for pastoralists, timber-use, biodiversity, livestock production. The origins can be
and ecosystem function. Changes in tree traced to the Drought Investigation Commis-
cover can have important ecosystem con- sion Report of 1923, in which many recom-
sequences with feedbacks to the earth- mendations pointed to the development
atmosphere system, including the role of of a focused attempt to improve veld and
trees as carbon sinks and their influence on livestock management and soil conserva-
energy budgets and hydrology. tion. At the time, vegetation management
in conservation areas was considered to
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