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et al. (2018), it would appear that in many circumstances providing a well-mixed ration that reduces the opportunity for diet selection and is fed ad libitum, may in many circumstances be as important as the composition of the diet or the particle length of the forage.


Conclusions There are many factors that influence rumen pH, including forage to concentrate ratio, forage particle length, dietary starch content, rate of starch release in the rumen and concentrate particle size, making an accurate prediction under commercial conditions problematic. This is further compounded by difficulties in accurately measuring rumen pH throughout the day on-farm. Daily saliva production is important in buffering the release of VFA, although dietary effects on saliva production are often smaller than anticipated. The particle size distribution of grass silage and grass silage based mixed rations in UK dairy herds is considerably longer than current North American guidelines for forages or rations. This suggests that the particle size of UK dairy rations is either too long, or that new guidelines or methods of determining particle size distribution are required. Feeding a short particle size grass silage or high levels of starchy concentrates may often have a smaller effect on rumen pH than anticipated, unless the diet contains high amounts of very finely ground starch or sugars sources, with a well-mixed ration that reduces the opportunity for selection being as important in many circumstances. However, on many UK dairy farms diets are poorly mixed and promote selection, which in combination with the restricted feeding that occurs on a high proportion, may be contributing to SARA.


Acknowledgements Much of the data presented in this paper was derived from the PhD thesis of Usama Tayyab. The author is also grateful to Prof. Chris Reynolds at Reading University, and Dr. Robert Wilkinson at Harper Adams University for their input into the studies, and AHDB Dairy for funding the work.


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Dijkstra, J., Ellis, J. L., Kebreab, E., Strathe, A. B., López, S., France, J. and Bannink, A. (2012). Ruminal pH regulation and nutritional consequences of low pH. Anim. Feed Sci. Tech. 172:22-33. Heinrichs, J., (2013). The Penn State Particle Separator. Retrieved on 19 July 2016 from http://extension.psu.edu/animals/dairy/nutrition/forages/ forage-quality-physical/separator/extension_publication_file Humer, E., Petri, R. M., Aschenbach, J. R., Bradford, B. J., Penner, G. B., Tafaj, M., Südekum, K. H. and Zebeli, Q. (2018). Practical feeding and management recommendations to mitigate the risk of subacute ruminal acidosis in dairy cattle. J. Dairy Sci. 101: 872-888. Jian, F. G., Lin, X. Y., Yan, Z. G., Hu, Z. Y., Liu, G. M., Sun, Y. D., Liu, W. and wang, Z. H. (2017). Effect of dietary roughage level on chewing activity, ruminal pH, and saliva secretion in lactating Holstein cows. J. Dairy Sci. 100: 2660-2671. Kmicikewycz, A. D., Harvatine, K. J. and Heinrichs, A. J. (2015). Effects of corn silage particle size, supplemental hay, and forage-to-concentrate ratio on rumen pH, feed preference, and milk fat profile of dairy cattle. J. Dairy Sci. 98:4850-4868. Maulfair, D. D., Zanton, G. I., Fustini, M. and Heinrichs, A.J. (2010). Effect of feed sorting on chewing behavior, production, and rumen fermentation in lactating dairy cows. J. Dairy Sci. 93:4791-4803. Mertens, D.R. (1997). Creating a system for meeting the fiber requirements of dairy cows. J. Dairy Sci. 80:1463-1481. Rojo-Gimeno, C., Fievez, V. and Wauters, E. (2018). The econonomic value of information provided by milk biomarkers under different scenarios: Case-study of an ex-ante analyis of fat-to-protein ration and fatty acid profile to detect subacute ruminal acidosis in dairy cows. Livestock Science 211: 30-41. Sinclair, K.D., Garnsworthy, P.C., Mann, G.E and Sinclair, L.A. (2014). Reducing dietary protein in dairy cow diets: implications for nitrogen utilization, milk production, welfare and fertility. Animal 8: 262-274. Tayyab, U. (2019). Evaluation of physically effective fibre in forages and its interaction with concentrate supplementation on rumen function, performance and health of UK dairy cows. A PhD thesis submitted to Harper Adams University, March 2019. Tayyab, U., Wilkinson, R. G., Charlton, G.L., Reynolds, C. K. and Sinclair, L. A. (2019). Grass silage particle size when fed with or without maize silage alters performance, reticular pH and metabolism of Holstein-Friesian dairy cows. Animal 13: 524-532. Tayyab, U., Wilkinson, R. G., Reynolds, C. K. and Sinclair, L. A. (2018). Particle size distribution of forages and mixed rations, and their relationship with ration variability and performance of UK dairy herds. Livestock Science, 217: 108-115. White, R. R., Hall, M. B., Firkins, J. L. and Kononoff, P. J. (2017). Physically adjusted neutral detergent fiber system for lactating dairy cows rations. I: Deriving equations that identify factors that influence effectiveness of fiber. J. Dairy Sci. 100 Zebeli, Q., Aschenbach, J.R., Tafaj, M., Boghun, J., Ametaj, B.N and Drochner, W. (2012). Review: Role of physically effective fiber and estimation of dietary fiber adequacy in high-producing dairy cattle. J. Dairy Sci. 95:1041-1056.


This article is based on the text of a paper presented by Liam Sinclair at the recent Ruminants Meeting of the Society of Feed Technologists


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