Mao et al.—New early Permian crinoid fauna 92(6):1066–1080


wider than long, length 9.4mm, width 15.5mm; articular facets plenary, subhorizontal broad, transverse ridge marked by fine crenulations (Fig. 5.1). Three anal plates in cup, large primanal (equivalent to the radianal) obliquely between two basals, mostly to left ofCradial, not in contact withDradial, supporting secundanal (equivalent to the anal X) and tertanal above (Fig.


5.3). Arms transversely well rounded, branching isotomously on first primibrachials; three thick first primibrachials (Fig. 5.2), length 6mm, width 13mm; secundibrachials cuneate in prox- imal portions, length 4mm, width 6.8; distal part of arms not preserved. Column circular, 6mm in diameter (Fig. 5.4).


Etymology.—The specific name is in honor of amateur collector Mr. Qiao Liman, Anyang, for help with fieldwork.


Material.—Holotype: NIGP 166470, only known specimen of this species.


Remarks.—The partial crown of U. qiaoi n. sp. is crushed flat. It differs from the most similar U. neverovoensis Mirantsev and Rozhnov, 2011 by having thicker plates and more impressed sutures. The new species also differs from U. grishini Mirantsev and Rozhnov, 2011 in the less elongate cup and relatively long infrabasal circlet and differs from U. karchevskyi Mirantsev and Rozhnov, 2011 in the strongly up-flared infrabasal circlet. It differs from all North American congeners in the presence of surface ornamentation.


Superfamily Erisocrinacea Wachsmuth and Springer, 1886 Family Erisocrinidae Wachsmuth and Springer, 1886 Genus Sinocrinus Tien, 1926


Type species.—Sinocrinus granulatus (Wanner, 1924).


Remarks.—Note Sinocrinus was erected based on Sinocrinus microgranulosus Tien, 1926, which is a junior subjective synonym of Sinocrinus granulatus Wanner, 1924 (cf. Sheffield, 2015).


Sinocrinus sheareri Strimple and Watkins, 1969 Figure 4.10–4.12


1969 Sinocrinus sheareri Strimple and Watkins, p. 182, pl. 39, figs. 8–11.


1976 Sinocrinus asymmetricus; Strimple, p. 636, fig. 3A–B. 1977 Sinocrinus sheareri; Webster, p. 156. 1986 Sinocrinus asymmetricus; Webster, p. 284. 2015 Sinocrinus sheareri; Sheffield, p. 468, figs. 7, 9A–C.


Occurrences.—Pennsylvanian, Atokan, Big Saline Formation, Soldiers Hole Member, United States, Texas; Pennsylvanian, Moscovian, Bashkirian? Spain; Taiyuan Formation, Dajian Member, early Permian (Asselian), Anyang, Henan, North China Block.


Description.—Dorsal cup flat, asymmetrically bowl-shaped (Fig. 4.12), length 6mm, width 21.4mm; plate ornamentation


1075


dense nodes, impressed sutures. Infrabasals five, fuse into a single plate, not in contact with radials; infrabasal circlet sub- horizontal, positioned slightly above basal plane, width 8.3mm, not as high as radial circlet; only basal and radial plates visible in side view. Basals five, tumid, unequal size with the smallest one in the posterior, the proximal portion of basals in the basal plane and the distal portion sharply upward to form part of lateral walls, length 6mm, width 6mm; interbasal sutures short. Radials tumid, equal size, the proximal tips of radials in basal plane of cup but not entering basal concavity, length 5.2mm, width 11.8mm; radial facets plenary (Fig. 4.11), outer ligament furrow poorly developed, sharply defined ligament pit, wide transverse ridges bearing fine denticles, well-developed oblique furrows, broad muscle scars, and small V-shaped intermuscular notch. No anal plate in external view, small internal anal notch on distal extremities of adjacent C and D radials in Figure 4.12. Column large, circular, with strongly crenulated cicatrix.


Material.—Figured specimen NIGP 166471.


Remarks.—Sheffield (2015) refined Sinocrinus Tien, 1926 using principal component analysis and neighborhood cluster analysis and extended the temporal span of the genus from the late Carboniferous (Moscovian) through the early Permian (Sakmarion). Sinocrinus asymmetricus Strimple, 1976 was rejected as the junior synonym of S. sheareri Strimple and Watkins, 1969 because of their qualitative and quantitative similarities with one another. Sinocrinus is mainly reported from the Houkou Limestone,


Taiyuan Series of northern China. This specimen is distin- guished from S. microgranulosus Tien, 1926, S. microgranu- losus pentalobus Tien, 1926, and S. linchenensis Tien, 1926 in having a proportionately lower, asymmetric dorsal cup, a proportionately larger column, lesser part of basals in cup walls, and short interbasal sutures. Three other forms are described by Tien (1926) as S. houkouensis, S. nodosus, and S. nodosus spinosus based on disarticulated plates. Sinocrinus sheareri has been reported from the Pennsylva-


nian of United States and Spain. One well-preserved specimen is assigned to S. sheareri, extending the stratigraphic range of S. sheareri to early Permian and the paleogeographic range to include China.


Genus Erisocrinus Meek and Worthen, 1865. Type species.—E. typus Meek and Worthen, 1865.


Erisocrinus cf. E. longwelli Lane and Webster, 1966 Figure 5.5, 5.6


Occurrence.—Taiyuan Formation,DajianMember, early Permian (Asselian), Anyang, Henan, North China Block.


Description.—Partial crown crushed flat, elongate cylinder (Fig. 5.5), length 52.7mm; dorsal cup large, medium bowl- shaped, length 13mm, width 33mm; plate ornament dense nodes. Infrabasals not preserved. Basals large, length 7.8mm, width 11mm, two preserved in Figure 5.5 and two visible in Figure 5.6. Radials large, length 8mm, width 14mm, swollen


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