Mao et al.—New early Permian crinoid fauna 92(6):1066–1080


Table 2. Measurements for Celonocrinus cf. C. expansus specimens. Units=mm.


Plates/specimens Length basal


Width basal Length radial Width radial


Length primibrachial 1 Width primibrachial 1 Length primibrachial 2 Width primibrachial 2


NIGP 166462 5.8


8.2 4.7


14 3 7


1.8 6.3


NIGP 166463 Material.—Holotype: NIGP 166464. 5.5


18 4 5 2


6.8 Measurements for plates of the specimens are given in Table 2.


Materials.—Figured specimens NIGP 166462 and NIGP 166463.


Remarks.—The two cups are not well preserved; NIGP 166462 (Fig. 3.1) retains one relatively complete radial, one basal, and 12 arm fragments, but only two fragmentized arms and radials of NIGP 166463 are preserved (Fig. 3.2, 3.3). Although not com- plete, the pentagonal basals, wide and short radials, radial facet in contact with both first primibrachial and first secundibrachials, wide and flat arms and flat brachials, and branching mode are very suggestive of Celonocrinus expansus.


Superfamily Erisocrinacea Wachsmuth and Springer, 1886


Family Erisocrinidae Wachsmuth and Springer, 1886


Genus Neoprotencrinus Knapp, 1969


Type species.—Paradelocrinus subplanus Moore and Plummer, 1940.


Neoprotencrinus anyangensis new species Figure 3.4


Holotype.—NIGP 166464.


Diagnosis.—Neoprotencrinus with radials not in basal circlet, primibrachials and six uniserial secundibrachials, and slightly impressed sutures.


Occurrence.—TaiyuanFormation,DajianMember,earlyPermian (Asselian), Anyang, Henan, North China Block.


Description.—Crown elongate cylindrical, length 85mm; cup low bowl shape, length 6.5mm, width 27mm; basal concavity very shallow to flat; plates smooth, sutures depressed. Basals not visible in side view; radials large, length 6.5mm, width 14mm, width commonly about twice the length. Proximal tips of radials not in the basal circlet; radial facets plenary. No anal plates visible with arms attached. Arms 10, moderately long, closely appressed; branching isotomously on first primibrachials; first primibrachials slightly spinose; primibrachials and six uniserial secundibrachials.


Remarks.—One flattened specimen is assigned to N. any- angensis. Only one side of holotype NIGP 166464 (Fig. 3.4) is preserved and is slightly crushed without anal plates. Large first secundibrachials are followed by five quadrate uniserial secun- dibrachials with the sixth uniserial secundibrachials with an angular distal suture for the reception of the first wedge-shaped brachial. Thereafter, the brachials are biserial. Webster and Kues (2006) moved Neoprotencrinus from the


Protencrinidae, transferring it to the Erisocrinidae because of its biserial arms. Neoprotencrinus are reported from the Pennsyl- vanian of the United States, including one species, N. brachiatus Moore and Plummer, 1940, with the arms preserved. The differences in shape of the radials and primibrachials along with the difference in the number of nonbiserial secundibrachials are the justification for the species. N. anyangensis is distinguished from N. brachiatus by the nonbiserial secundibrachials, from N. gutschicki Webster and Kues, 2006 by more impressed sutures, and from Neoprotencrinus cranei Strimple, 1949, N. regulates Strimple, 1949, N. disculus Strimple, 1949, and N. subplanus rockensis Moore and Plummer, 1940 by proximal tips of radials not in the basal circlet.


Family Protencrinidae Knapp, 1969 Genus Protencrinus Jaekel, 1918


Type species.—P. moscoviensis Jaekel, 1918.


Protencrinus baliensis Webster, 2012b Figure 4.1–4.3


Occurrences.—Early Permian, series undesignated, Crete, Greece; Taiyuan Formation, Dajian Member, early Permian (Asselian), Anyang, Henan, North China Block.


Description.—Cup flat bowl shape (Fig. 4.3), medium-sized basal concavity, pentagonal in oral outline, smooth plate sculpturing, length 5mm, width 23mm. Infrabasals five, pen- tagonal, truncated distally in contact with radials, equal size, longer than wide, length 5mm, width 4mm; infrabasal circlet confined to basal concavity, down-flaring. Basals five, small quadrangular, not in lateral contact (Fig. 4.1), length 3mm,


width 3mm, proximal tips in basal concavity, distal tips up- flaring with tips barely visible in lateral view. Radials five, hexagonal, much wider than long, length 5mm, width 14mm, gently convex transversely, moderately convex longitudinally, proximal tips in basal concavity, distally up-flaring form lateral walls; radial facet plenary (Fig. 4.2), deep ligament pit; narrow outer marginal area; deep central pit and intermuscular furrow; muscle fields shallowly concave; narrow lateral ridges. Anal notch not visible in lateral view with a narrow distinct V-shaped groove on the internal edge (Fig. 4.3). Column facet circular, 4mm in diameter, impressed in infrabasal circlet.


Material.—Figured specimen NIGP 166465.


1071


Etymology.—Named after Anyang city, Henan Province, which hosts the type locality.


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