Journal of Paleontology, 92(6), 2018, p. 1066–1080 Copyright © 2018, The Paleontological Society 0022-3360/15/0088-0906 doi: 10.1017/jpa.2018.27


A new crinoid fauna from the Taiyuan Formation (early Permian) of Henan, North China


Yingyan Mao,1,2,3 Gary D. Webster,4 William I. Ausich,5 Yue Li,1 Qiulai Wang,1,2 and Mike Reich3,6,7


1Key Laboratory of Economic Stratigraphy and Paleogeography, Nanjing Institute of Geology and Paleontology, Chinese Academy of Sciences, Nanjing 210008, China ⟨yymao@nigpas.ac.cn⟩, ⟨yueli@nigpas.ac.cn⟩, ⟨qlwang@nigpas.ac.cn⟩ 2University of the Chinese Academy of Sciences, Beijing 100049, China 3SNSB - Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Straße 10, 80333 Munich, Germany ⟨m.yingyan@lrz.uni-


muenchen.de⟩, ⟨m.reich@lrz.uni-muenchen.de⟩ 4School of Environmental Sciences, Washington State University, Pullman, Washington, USA ⟨webster@wsu.edu⟩ 5School of Earth Sciences, The Ohio State University, Columbus, Ohio 43210, USA ⟨ausich.1@osu.edu⟩ 6Department of Earth and Environmental Sciences, Division of Palaeontology and Geobiology, Ludwig-Maximilians-Universität München,


Richard-Wagner-Strasse 10, 80333 München, Germany ⟨mike.reich@lmu.de⟩ 7GeoBio-Center, Ludwig-Maximilians-Universität München, Richard-Wagner-Straße 10, 80333 Munich, Germany


Abstract.—A diverse Permian crinoid fauna is reported from the Taiyuan Formation, Dajian Member (Asselian) at Anyang, northeastern Henan Province of the North China Craton. The specimens are well preserved, including articu- lated crowns and cups. The fauna contains representatives of each of the major Paleozoic crinoid clades: Cladida (including the Flexibilia), Disparida, and Camerata. Identified genera suggest a greater affinity with North American faunas than with Tethyan faunas. Four new species, Neoprotencrinus anyangensis, Ulocrinus qiaoi, Artichthyocrinus limani,and Synbathocrinus chenae, are proposed herein.


UUID: http://zoobank.org/D08DDDCD-485C-45CC-A014-C1CB58C26588 Introduction


Permian crinoids are relatively rare compared with crinoids from earlier parts of the Paleozoic (Table 1). The most diverse and abundant articulated Permian crinoid crowns and cups were reported from Timor (Wanner, 1916, 1924, 1937, among others; Webster, 2012a; Webster and Donovan, 2012). Although the stratigraphic position was uncertain, it was considered to be Artinskian by Charlton et al. (2002). In addition, early Permian crinoid faunas are known from Russia (Yakovlev, 1926, 1927, 1930; Yakovlev and Ivanov, 1956), the United States (the midcontinent: Moore and Plummer, 1940; Pabian and Strimple, 1974; southern Nevada: Lane and Webster, 1966; Webster and Lane, 1967, 2007), Australia (eastern:Willink, 1978, 1979a, b, 1980; Webster and Jell, 1999b; Western Australia: Webster, 1987, 1990; Webster and Jell, 1992; Teichert and Webster, 1993; Tasmania: Sieverts-Doreck, 1942), Oman (Webster and Sevastopulo, 2007; Webster et al., 2009b), Thailand (Webster and Jell, 1993), British Columbia (Webster et al., 2009a), Greece (Webster, 2012b), Bolivia (Branisa, 1965; Strimple and Moore, 1971; Burke and Pabian, 1978), Mexico (Strimple, 1971), China (Tien, 1926; Chen and Yao, 1993), and Spitsber- gen/Norway (Gorzelak et al., 2013). Crinoid faunas are also known from the middle Permian of Tunisia (Valette, 1934; Lane, 1979), Europe (Gregorio, 1930; Ramovš and Sieverts- Doreck, 1968; Strimple and Sevastopulo, 1982), and Pakistan (Waagen, 1887), as well as from upper Permian strata of Europe


(e.g., Donovan et al., 1986; Reich, 2007). Among them, the Timor, southern Nevada, Western Australia, and southern Ural Mountains faunas are the four main early Permian faunas representing more than 40 genera. The only Permian crinoids reported previously from China are Separocrinus discoides Chen and Yao, 1993 (Webster et al., 2009c) from the early Permian Sakmarian to Artinskian of Yunnan and one possible crinoid fauna from North China (southern Hebei Province), Taiyuan Series (exact age uncertain), including Delocrinus, Mathericrinus, and Sinocrinus (Tien, 1926). The fossils described herein were collected from an outcrop


at Tianxi Village, Shanying Town, southern Anyang City, Henan Province (36.24730°N, 114.65929°E) (Fig. 1), approxi- mately 120km south of the Lincheng Coal Field Section of southern Hebei Province studied by Tien (1926). The present study adds to our knowledge of the diversity of crinoids during the earliest Permian in the North China Block.


Geologic setting and stratigraphy


Lower Paleozoic strata of the eastern part of the North China Block (or Sino-Korean Platform) are composed dominantly of shallow marine carbonates (Feng et al., 1990). The block was largely uplifted during Ordovician time. A remarkable sedi- mentary gap (parallel unconformably) is present at the top of the Cambrian or Ordovician. The dividing line is from Xiaxian to Dengfeng. In the south of the dividing line, the gap is at the top


1066


Page 1  |  Page 2  |  Page 3  |  Page 4  |  Page 5  |  Page 6  |  Page 7  |  Page 8  |  Page 9  |  Page 10  |  Page 11  |  Page 12  |  Page 13  |  Page 14  |  Page 15  |  Page 16  |  Page 17  |  Page 18  |  Page 19  |  Page 20  |  Page 21  |  Page 22  |  Page 23  |  Page 24  |  Page 25  |  Page 26  |  Page 27  |  Page 28  |  Page 29  |  Page 30  |  Page 31  |  Page 32  |  Page 33  |  Page 34  |  Page 35  |  Page 36  |  Page 37  |  Page 38  |  Page 39  |  Page 40  |  Page 41  |  Page 42  |  Page 43  |  Page 44  |  Page 45  |  Page 46  |  Page 47  |  Page 48  |  Page 49  |  Page 50  |  Page 51  |  Page 52  |  Page 53  |  Page 54  |  Page 55  |  Page 56  |  Page 57  |  Page 58  |  Page 59  |  Page 60  |  Page 61  |  Page 62  |  Page 63  |  Page 64  |  Page 65  |  Page 66  |  Page 67  |  Page 68  |  Page 69  |  Page 70  |  Page 71  |  Page 72  |  Page 73  |  Page 74  |  Page 75  |  Page 76  |  Page 77  |  Page 78  |  Page 79  |  Page 80  |  Page 81  |  Page 82  |  Page 83  |  Page 84  |  Page 85  |  Page 86  |  Page 87  |  Page 88  |  Page 89  |  Page 90  |  Page 91  |  Page 92  |  Page 93  |  Page 94  |  Page 95  |  Page 96  |  Page 97  |  Page 98  |  Page 99  |  Page 100  |  Page 101  |  Page 102  |  Page 103  |  Page 104  |  Page 105  |  Page 106  |  Page 107  |  Page 108  |  Page 109  |  Page 110  |  Page 111  |  Page 112  |  Page 113  |  Page 114  |  Page 115  |  Page 116  |  Page 117  |  Page 118  |  Page 119  |  Page 120  |  Page 121  |  Page 122  |  Page 123  |  Page 124  |  Page 125  |  Page 126  |  Page 127  |  Page 128  |  Page 129  |  Page 130  |  Page 131  |  Page 132  |  Page 133  |  Page 134  |  Page 135  |  Page 136  |  Page 137  |  Page 138  |  Page 139  |  Page 140  |  Page 141  |  Page 142  |  Page 143  |  Page 144  |  Page 145  |  Page 146  |  Page 147  |  Page 148  |  Page 149  |  Page 150  |  Page 151  |  Page 152  |  Page 153  |  Page 154  |  Page 155  |  Page 156  |  Page 157  |  Page 158  |  Page 159  |  Page 160  |  Page 161  |  Page 162  |  Page 163  |  Page 164  |  Page 165  |  Page 166  |  Page 167  |  Page 168  |  Page 169  |  Page 170  |  Page 171  |  Page 172  |  Page 173  |  Page 174  |  Page 175  |  Page 176  |  Page 177  |  Page 178  |  Page 179  |  Page 180  |  Page 181  |  Page 182  |  Page 183  |  Page 184  |  Page 185  |  Page 186  |  Page 187  |  Page 188  |  Page 189  |  Page 190