Huang et al.—New selenothemistid genus from China 92(6):1049–1053
anterior; RP=radius posterior; sd=subdiscoidal space; Sn= subnodal crossvein. All measurements are given in millimeters.
Repositories and institutional abbreviations.—NIGP: Nanjing Institute of Geology and Palaeontology CAS, Nanjing, China.
Systematic paleontology
Superorder Odonatoptera Martynov, 1932 Order Odonata Fabricius, 1793 Clade Isophlebioptera Bechly, 1996
Family Selenothemistidae Handlirsch, 1939
Type genus.—Selenothemis Handlirsch, 1920. Other genera: Turanothemis Pritykina, 1968, Sinothemis new genus, Car- aphlebia Carpenter, 1969, Paraliassophlebia Hong, 1982.
Genus Sinothemis new genus Type species.—Sinothemis difficilis n. sp.
Etymology.—Named after Sinica, Latin name for China, and the suffix Themis (ancient Greek Titaness, personification of the divine order), frequently employed for taxa in this family. Gender masculine.
Remark.—Sinothemis shares some characters with Selenothe- mis and others with Turanothemis. Its exact affinities will need a phylogenetic analysis to be solved.
Sinothemis difficilis new species Figures 2–4
Holotype.—NIGP167785, stored at the Nanjing Institute of Geology and Palaeontology CAS, Nanjing, China.
Diagnosis (for genus and species).—Hindwing characters only. Wing ~ 41mm long; base of RP2 not aligned with subnodus; only one oblique vein ‘O’; only one row of cells between RP3/4
Figure 3. Sinothemis difficilis n. gen. n. sp., holotype NIGP167785, reconstruction of hindwing. Ax=primary antenodal crossvein; CuA=cubitus anterior; d=discoidal cell; IR=intercalary radial veins; MAa=median anterior; MP=median posterior; N=nodus; Pt=pterostigma; RP=radius posterior; sd=subdiscoidal space; Sn=subnodal crossvein. Scale bar=8mm.
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and MAa well distal (five cells) of nodus level; only one row of cells between IR2 and RP3/4 well distal (three cells) of oblique vein.
Occurrence.—Guancaishan, Jianping County, Liaoning Pro- vince, NE China; late Oxfordian to early Kimmeridgian.
Description.—A thorax with nearly complete abdomen, frag- ments of legs, a nearly complete hindwing plus fragments of another attached. Hindwing probably hyaline, 40.8mmlong, 10.0mmwide
at nodus; distance between base and arculus 5.4mm, between arculus and nodus 11.7mm, between nodus and pterostigma 15.0mm, between pterostigma and apex 5.8mm; wing very shortly petiolate; cells of anal area irregular, some being very small and others being very large, a large cell below CuP; median space free; submedian space with a curved vein CuP; a curved crossvein separating submedian and subdiscoidal spaces; subdiscoidal space posteriorly closed and very broad, with AA making a right angle in its basal part; discoidal cell free, basally closed, basal side 0.7mmlong, costal side 1.2mm long, posterior side 1.5mm long, distal side 2.0mm long; RP
Figure 2. Sinothemis difficilis n. gen. n. sp., holotype NIGP167785, photograph of general habitus. Scale bar=10mm.
Figure 4. Sinothemis difficilis n. gen. n. sp., SEM scans showing the details on body surface (pattern and setae). (1) Detail of anterior part of third segment; (2) posterior part of first segment and anterior part of second; (3) detail of anterior part of first segment; (4) detail of anterior part of fifth segment.
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